Manota capillata, Hippa, Heikki & Ševċík, Jan, 2010

Hippa, Heikki & Ševċík, Jan, 2010, Notes on Oriental and Australasian Manotinae (Diptera, Mycetophilidae), with the description of thirteen new species, Zootaxa 2333, pp. 1-25 : 7-8

Manota capillata sp. n.

Figs 5 A, B, C

Male. Colour. Both specimens seem strongly faded. Head pale brown, frons, vertex and part of occiput darker brown. Antenna unicolorous pale brown. Mouthparts pale yellowish. Thorax brownish, darker than legs. Legs pale yellowish, apical third of femur 3 slightly infuscated. Wing unicolorous pale yellowish-brown. Haltere yellowish brown, knob blackish (blackish granules inside). Abdomen (treated with KOH) pale brown. All setosity pale, yellowish or brownish, thicker setae appearing darker than finer setae and trichia. Head. Antennal flagellomere 4, Fig. 5 A. Palpomere 3 of maxillary palpus with apicomesial extension, with 3–4 apically expanded curved sensilla; palpomere 4 with parasegment; palpomere 5 ca.1.1 times as long as palpomere 4. Nine to eleven strong postocular setae. Thorax. Anepisternum setose, with 21–35 setae; anterior basalare non-setose; preepisternum 2 setose, with 15–16 setae; laterotergite non-setose; episternum 3 setose, with 9–12 setae. Wing. Similar to Fig 1 A; wing length 2.0 mm. Hypopygium, Figs 5 B, C. Sternite 9 short, less than half ventral length of gonocoxa, laterally fused with gonocoxa, posterior margin convex, anterior margin nearly straight, without any indication of medial incision, setae similar to ventral setae of gonocoxa. Ventral mesial margin of gonocoxa simple, basal part united with anterior part of parastylar lobe, the latter with several setae which are similar to gonocoxal setae. Paraapodemal lobe a curved sclerotized stripe ( Fig. 5 B), in holotype concealed under mesial margin of gonocoxa in ventral view, in paratype partly exposed. Dorsal mesial margin of gonocoxa indistinct, fused with a large crescent-shaped lobe bearing a sub-marginal zone of long setae on its ventral side. Gonocoxa without an apico-dorsal lobe or apophysis. Two juxtagonostylar setae present: a curved megaseta and a much smaller similarly curved normal seta, both arising from a common low basal body. Gonostylus short, transverse, narrowing apically, apico-ventral part forming an indistinct lobe, setae on ventral side moderately long, on apico-ventral lobe longer, in a row on posterior margin, dorsal side nonsetose. Aedeagus elongate subtriangular, rather broad apically, constricted basally, the lateral shoulders distinct, apex curved ventrad. Hypoproct large, posteriorly extending as far as gonostylus, ventrally with ca. 40 scattered setae on each half. Cerci mesially separate.

Female unknown.

Discussion. M. capillata is similar to M. parvistylata Hippa known from the Malay Peninsula. It is distinguished e.g. by having few and short setae at the postero-ventral and posterolateral margin of the gonocoxa, similar to the more anteriorly placed setae, while in M. parvistylata these setae are very long, the longest ones more than twice the length of the more anteriorly placed setae and appear as a more or less distinct transverse row. In M. capillata the setae on the dorsomesial lobe of the gonocoxa are longer, as long as or longer than the juxtagonostylar megaseta, whereas in M. parvistylata they are shorter than the megaseta. In both mounts of M. capillata the gonostylus appears similar, with broad base and narrow apex, and rather different from the more parallel sided one in the holotype of M. parvistylata (Hippa 2008) . In both species the gonostylus is very similar, but its position in a mount is very critical.

Etymology. The name is from the Latin, capillata , hairy, and refers to the long hairs on the dorsal lobe on the gonocoxa.

Types. Holotype. Male, INDONESIA, Sumatra, Sumatera Utara, Semangat Gunung, 1300 m, Malaise in jungle, 16.03.1992, H. Hippa (in SMNH).

Paratypes. 1 male with same data as holotype (in SMNH).

Figures

FIGURE 1. Wing, dorsal view (A, B, C, E) and basal part of wing, ventral view (D). A. Manota bruneiensis sp. n. (paratype) B. M. subforceps sp. n. (holotype). C. M. sinepollex sp. n. (holotype). D, E. Eumanota vilkamaai sp. n. (holotype). Scale 1.0 mm.

FIGURE 5. Manota capillata sp. n. (holotype). A. Antennal flagellomere 4, lateral view. B. Hypopygium, dorsal view. C. Hypopygium, ventral view. Scale 0.10 mm.