Maurhinophora Cerretti & Pape, 2020

Cerretti, Pierfilippo, Badano, Davide, Gisondi, Silvia, Giudice, Giuseppe Lo & Pape, Thomas, 2020, The world woodlouse flies (Diptera, Rhinophoridae), ZooKeys 903, pp. 1-130 : 1

Maurhinophora Cerretti & Pape gen. nov. Fig. 14

Unambiguous character state changes

(Table 1 , Fig. 20 ). Global apomorphies: none; local apomorphies: 13:1, 14:1, 21:2; 30:2, 42:1, 44:1.

Diagnosis.

Head: head higher than long in lateral view. Facial ridge 1.1 times as long as frons. Ocellar setae virtually absent. Frons approx. 0.9 times as wide as compound eye in dorsal view. Median (= inner) vertical setae strong and crossed. Five medially crossed frontal setae, slightly reclinate, descending to approx. half level of pedicel. Fronto-orbital plate with some short setulae. Two proclinate orbital setae. One upper lateroclinate orbital seta. Parafacial bare, at its narrowest point at most 1.5 times as wide as maximum diameter of arista. Vibrissal angle receding. Vibrissa well developed, arising slightly below level of lower facial margin. Lower facial margin sunken and not visible in lateral view. Facial ridge slightly and evenly convex with a row of setae on lower 2/3, decreasing in size dorsally. Face deeply concave, antennae hidden in lateral view. Antenna long and narrow, much longer than height of gena. Postpedicel narrowly elongated approx. 5 times as long as pedicel. Arista bare (or apparently so). Arista thickened in proximal 2/5-1/2; second aristomere at most as long as wide. Lunule bare. Gena, in profile, approx. 1/5 as high as compound eye. Palpus absent.

Thorax: prosternum bare. Postpronotum with three setae arranged in triangle. Three postsutural supra-alar setae (first postsutural supra-alar seta well developed, i.e., longer than posterior notopleural seta and approx. the same size as anterior notopleural seta). Scutellum with one pair of well-developed basal setae and one pair of strong, horizontal and crossed apical setae; basal setae placed at level of apical setae. Anatergite with a tuft of short setulae below lower calypter. Subscutellum moderately swollen, not fully sclerotised. Posterior lappet of metathoracic spiracle larger than anterior lappet (as in Baniassa ). Lower calypter distinctly tongue-shaped (ground-plan trait of Rhinophoridae ) (Fig. 2E ). Costal sector cs2 setose ventrally. Costal spine not differentiated from general costal setae. Costal sector cs5 clearly shorter than costal sector cs2. Vein R1 entirely setulose dorsally. Vein R4+5 with setulae dorsally extending from base to approx. level of bend of vein M1. Bend of vein M1 well developed, rounded and well removed from wing margin. Crossvein dm-m forming a right angle with proximal section of M4. Vein CuA+CuP not reaching wing margin. Preapical anterodorsal seta of fore tibia longer than preapical dorsal seta. Fore tarsus not compressed. Tibiae of mid and hind leg normally developed. Mid tibia with one, short subdistal anterodorsal seta. Hind tibia with 3 dorsal preapical setae.

Abdomen: tergites without microtomentum and with relatively strong and suberect general setulae; syntergite 1+2 without median discal setae, tergite 3 with one pair of strong median discal setae, tergites 4 and 5 with a row of strong marginal setae (discal setae not differentiated).

Distribution.

Afrotropical - Mauritius.

Type species.

Maurhinophora indoceanica Cerretti & Pape, sp. nov., by present designation.

Etymology.

The generic name is a composite word formed from the first part of the name of the island Mauritius, to which the known species is restricted, and the name Rhinophora , which is the type-genus for the family-group name Rhinophoridae . The name should be treated as a feminine noun.

Figures

Figure 2. Morphological terminology of Rhinophoridae, head, thorax and abdomen. A Chaetotaxy of head, thorax and abdomen in dorsal view B, C arrangement of setae of postpronotum: B three setae arranged in shallow triangle C three setae standing in a nearly right-angled triangle D thorax, sclerites and chaetotaxy, in lateral view E wing veins and cells. Drawings by Giulia Bellanti.

Figure 14. Maurhinophora indoceanica gen. et sp. nov. ♀ (Mauritius) [holotype] A habitus in lateral view B abdomen in dorsal view C wing in dorsal view D head in lateral view.

Figure 20. Phylogeny of Rhinophoridae based on morphological evidence. Single tree obtained under IW (total fit: 62.33), enforcing a k-value of 4, with unambiguous character state changes mapped above branches [black squares: global apomorphies (i. e., uncontradicted and unreversed apomorphic character states; white squares: local apomorphies (i. e., homoplasious character state due to convergence or reversal)] and new taxa highlighted in bold.