Hypatopa

Adamski, David, 2013, Review of the Blastobasinae of Costa Rica (Lepidoptera: Gelechioidea: Blastobasidae), Zootaxa 3618 (1), pp. 1-223 : 77-79

Key to the Species of Hypatopa of Costa Rica

1. Male................................................................................................ 2

- Female............................................................................................. 50

2. Gnathos present)...................................................................................... 3

- Gnathos absent).......................................................................... Hypatopa rabio

3. Gnathos emarginate mesially ( Figs. 139 , 145 , 149, 151 , 153, 157 , 159, 161, 163 , 165, 167, 169 , 171 , 177, 179, 181 , 183, 185 , 189, 191 , 195 , 201, 203, 205 , 209, 211 , 213, 215 , 219, 221, 223 , 225 , 231 )........................................ 4

- Gnathos not emarginate mesially ( Figs. 137 , 147 , 173 , 141, 143 , 155 , 175 , 187 , 193 , 197, 199 , 207 , 217 , 227 , 231, 233 )... 36

4. Mesial emargination on a slightly raised or protuberant lobe ( Figs. 139 , 145 , 153 , 163 , 171 , 183 , 189 , 203, 205 , 209 , 221 , 225 , 231 )................................................................................................ 5

- Mesial emargination on an entire margin ( Figs. 149, 151 , 157 , 159, 161 , 167, 169 , 177, 179, 181 , 185 )................ 17

5. Apical process of ventral part of valva acutely curved in area from base to 1/2 ( Figs. 139 , 163 , 209 , 231 )................ 6

- Apical process of ventral part of valva not acutely curved in area from base to 1/2 ( Figs. 145 , 153 , 171 , 183 , 189 , 203, 205 , 221 , 225 )............................................................................................ 9

6. Apical process of ventral part of valva acutely curved laterally ( Figs. 231 )............................. Hypatopa rudis

- Apical process of ventral part of valva not acutely curved laterally (139, 163, 209).................................. 7

7. Proximal flange of valva longer than wide ( Fig. 163 ); anellus rounded apically ( Fig. 164 )............... Hypatopa plebis

- Proximal flange of valva not longer than wide ( Figs. 139 , 209 ); anellus emarginate apically ( Figs. 140 , 210 )............. 8

8. Gnathos deeply emarginate mesially; proximal flange wider than long ( Fig. 139 )...................... Hypatopa cladis

- Gnathos shallowly emarginate mesially; proximal flange about as wide as long ( Fig. 209 ).................. Hypatopa io

9. Margin of proximal flange of valva crenulate ( Figs. 183 , 225 )................................................. 10

- Margin of proximal flange of valva serrate ( Figs. 145 , 153 , 171 , 189 , 203, 205 , 221 )............................... 11

10. Gnathos deeply emarginate mesially ( Fig. 183 ); anellus truncate apically ( Fig. 184 )................... Hypatopa semela

- Gnathos shallowly emarginate mesially ( Fig. 225 ); anellus deeply emarginate, forming two lateral lobes and with two porrect median spinelike processes ( Fig. 226 )......................................................... Hypatopa solea

11. Gnathos deeply emarginate mesially ( Figs. 153 , 171 )........................................................ 12

- Gnathos shallowly emarginate mesially ( Figs. 145 , 189 , 203, 205 , 221 ).......................................... 13

12. Sclerite of phallus acutely curved near middle; vesica with barbed cornuti ( Fig. 154 )..................... Hypatopa crux

- Sclerite of phallus shallowly curved near middle; vesica without cornuti ( Fig. 172 ).................... Hypatopa caepae

13. Phallus and sclerite of phallus longer than valva ( Figs. 145 , 204, 206 )........................................... 14

- Phallus and sclerite of phallus shorter than valva ( Figs. 190 , 222 )............................................... 16

14. Sclerite of phallus acutely sigmoid-shaped ( Fig. 204 )............................................... Hypatopa dux

- Sclerite of phallus not acutely curved throughout length ( Figs. 146 , 206 )......................................... 15

15. Juxta divided mesially ( Fig. 145 ); anellus nearly parallelsided throughout most of length ( Fig. 146 )........ Hypatopa cotis

- Juxta bandlike ( Fig. 205 ); anellus constricted beyond base ( Fig. 206 )................................. Hypatopa erato

16. Gnathos widely emarginate ( Fig. 189 ); anellus near parallelsided throughout most of length ( Fig. 190 )...... Hypatopa rego

- Gnathos narrowly emarginate ( Fig. 221 ); anellus gradually narrowed from base ( Fig. 222 )............... Hypatopa umbra

17. Gnathos projected anteriorly near or greater than 1/2 width ( Figs. 169 , 177 , 191 , 195 , 201 , 219, 223 ).................. 18

- Gnathos not projected anteriorly as above ( Figs. 149, 151 , 157 , 159, 161 , 165, 167 , 179, 181 , 185 , 211 , 213, 215 )........ 24

18. Ventral part of valva greatly protracted inwards basally ( Figs. 169 , 195 , 23 )...................................... 19

- Ventral part of valva slightly protracted inwards basally ( Figs. 177 , 191 , 201 , 19 ).................................. 21

19. Proximal flange of valva nearly twice as long as wide; apical process of ventral part of valva acutely curved basally ( Fig. 169 )....................................................................................... Hypatopa manus

- Proximal flange of valva about as long as wide; apical process of ventral part of valva not acutely curved from base ( Figs. 195 , 223 )............................................................................................... 20

20. Apical margin of proximal flange wider than base ( Fig. 223 ); anellus widely emarginate, forming two lateral lobes, and with two convergent apical processes ( Fig. 224 )...................................................... Hypatopa texo

- Apical margin of proximal flange narrower than base ( Fig. 195 ); anellus rounded apically ( Fig. 196 )........ Hypatopa verax

21. Gnathos deeply emarginate mesially ( Figs. 177 , 191 , 219 ).................................................... 22

- Gnathos shallowly emarginate mesially ( Fig. 201 ).................................................. Hypatopa nox

22. Sclerite of phallus recurved apically ( Figs. 192 , 220 )......................................................... 23

- Sclerite of phallus not recurved apically ( Fig. 178 ).............................................. Hypatopa scobis

23. Margin of proximal flange serrate ( Fig. 191 ); sclerite of phallus recurved to near base of anellus ( Fig. 220 ).... Hypatopa ira

- Margin of proximal flange entire ( Fig. 219 ); sclerite of phallus recurved to subapical region ( Fig. 192 )...... Hypatopa styga

24. Ventrolateral margin of proximal flange with two subequal, large spinelike processes ( Fig. 149 ).......... Hypatopa hecate

- Ventrolateral margin of proximal flange entire, crenulate, or serrate ( Figs. 151 , 157 , 159, 161 , 165, 167 , 179, 181 , 185 , 211 , 213, 215 )........................................................................................... 25

25. Margin of proximal flange entire or crenulate.............................................................. 26

- Margin of proximal flange serrate........................................................................ 30

26. Base of phallus widely bulbous basally ( Figs. 152 , 180 , 214 ).................................................. 27

– Base of phallus not widely bulbous basally ( Figs. 160 , 166 )................................................... 29

27. Vesica with cornuti ( Fig. 152 )................................................................. Hypatopa acus

- Vesica without cornuti ( Fig. 180 , 214 ).................................................................... 28

28. Proximal flange elongate ( Fig. 213 ); sclerite of phallus straight basally ( Fig. 214 )......................... Hypatopa vox

- Proximal flange ellipsoid ( Fig. 179 ); sclerite of phallus acutely curved basally ( Fig. 180 )................. Hypatopa agnae

29. Anellus 1/2 length of phallus, nearly parallelsided ( Fig. 166 )....................................... Hypatopa dolo

- Anellus less than 1/2 length of phallus, wider on basal 2/3, abruptly narrowed apically ( Fig. 160 )......... Hypatopa arxcis

30. Juxta divided mesially ( Figs. 161 , 211 ).................................................................... 31

- Juxta bandlike ( Figs. 157 , 167 , 181 , 185 , 215 ).............................................................. 32

31. Apical process of ventral part of valva downcurved ( Fig. 161 ); anellus broadly rounded apically ( Fig. 162 ).... Hypatopa caedis

- Apical process of ventral part of valva upturned ( Fig. 211 ); anellus broadly emarginate apically ( Fig. 212 ).... Hypatopa leda

32. Anellus apically emarginate ( Figs. 158 , 182 , 216 )........................................................... 33

- Anellus apically rounded ( Figs. 168 , 186 )................................................................. 35

33. Anellus shallowly emarginate apically, bulbous basally ( Fig. 158 )................................... Hypatopa hera

- Anellus deeply emarginate apically, not bulbous basally ( Figs. 182 , 216 )......................................... 34

34. Emargination of anellus forming two apically rounded lateral lobes ( Fig. 182 )........................ Hypatopa phoebe

- Emargination of anellus forming two convergent, apically pointed lateral lobes ( Fig. 216 )............ Hypatopa tapadulcea

35. Proximal flange subrectangular; spinelike process of ventral part of valva rounded apically ( Fig. 167 )....... Hypatopa cyane

- Proximal flange semicircular; apical process of ventral part of valva acute ( Fig. 185 )..................... Hypatopa edax

36. Gnathos with a small or protuberant mesial lobe ( Figs. 137 , 147 , 155 , 173 , 187 )................................... 37

- Gnathos with mesial lobe absent ( Figs. 141, 143 , 175 , 193 , 197, 199 , 207 , 217 , 227 , 233 )........................... 41

37. Proximal flange of valva with 1–4 spinelike processes on apicoventral region ( Figs. 148 , 174 )........................ 38

- Proximal flange of valva without such processes ( Figs. 137 , 155 , 187 )........................................... 39

38. Proximal flange of valva elongate, densely setose ( Fig. 173 ); sclerite of phallus broadly curved ( Fig. 174 ).. Hypatopa cotytto

- Proximal flange of valva not elongate, sparsely setose ( Fig. 147 ); sclerite of phallus acutely curved at 2/3 ( Fig. 148 )................................................................................................. Hypatopa pica

39. Margin of proximal flange of valva entire ( Fig. 137 ); anellus with apical barb ( Fig. 138 )................... Hypatopa nex

- Margin of proximal flange serrate ( Figs. 155 , 187 ); anellus without apical barb ( Figs. 156 , 188 )...................... 40

40. Proximal flange serrate ( Fig. 155 ); apical margin of anellus narrower than base ( Fig. 156 )................ Hypatopa limae

- Proximal flange deeply serrate ( Fig. 187 ); anellus truncate apically ( Fig. 188 )........................ Hypatopa joniella

41. Apical process of ventral part of valva diminutive ( Figs. 141, 143 ); apex of anellus bearing a pair of recurrent lateral barblike processes ( Figs. 142, 144 ).............................................................................. 42

- Apical process of ventral part of valva of normal length ( Figs. 197, 199 , 207 , 217 , 227 , 233 ); anellus not bearing recurrent barblike processes ( Figs. 198, 200 , 208 , 218 , 228 , 234 )....................................................... 43

42. Anellus with crenulate barblike processes ( Fig. 142 ).............................................. Hypatopa juno

- Anellus with straight barblike processes ( Fig. 144 )............................................... Hypatopa actes

43. Anellus mesially emarginate apically ( Figs. 228 , 234 )........................................................ 44

- Anellus not mesially emarginate apically ( Figs. 176 , 194 , 198, 200 , 208 , 218 )..................................... 45

44. Anellus marginate, forming two large, divergent apicolateral processes ( Fig. 228 )................... Hypatopa bilobata

- Anellus marginate, forming two short apicolateral processes ( Fig. 234 )............................... Hypatopa musa

45. Phallus widely bulbous basally ( Figs. 194 , 198, 200 )......................................................... 46

- Phallus not widely bulbous ( Figs. 176 , 208 , 218 )............................................................ 48

46. Apical part of sclerite of phallus slightly curved ( Fig. 194 ).......................................... Hypatopa texla

- Apical part of sclerite of phallus recurved ( Figs. 198, 200 ).................................................... 47

47. Basal ridge of digitate process of dorsal part of valva densely setose; base of apical process of ventral part of valva with a set- ose lobe ( Fig. 199 )......................................................................... Hypatopa mora

- Basal ridge of digitate process of dorsal part of valva sparsely setose; base of apical process of ventral part of valva without a setose lobe ( Fig. 197 ).................................................................. Hypatopa tapadulcea

48. Apical process of ventral part of valva downcurved ( Fig. 217 )...................................... Hypatopa dicax

- Apical process of ventral part of valva not downcurved ( Figs. 175 , 207 )......................................... 49

49. Margin of proximal flange entire ( Fig. 175 ); sclerite of phallus shorter than phallus ( Fig. 176 )............ Hypatopa lucina

- Margin of proximal flange serrate ( Fig. 207 ); sclerite of phallus as long as phallus ( Fig. 208 )................ Hypatopa fio

50. Eighth tergum with a darkly pigmented median longitudinal streak ( Figs. 265 , 270 , 272–273 )........................ 51

- Eighth tergum without a darkly pigmented median longitudinal streak ( Figs. 261–264 , 266–269, 271 ).................. 54

51. Ductus bursae coiled throughout most of length ( Fig.265 )...................................... Hypatopa tapadulcea

- Ductus bursae not coiled throughout most of length ( Figs. 270 , 272–273 )........................................ 52

52. Posterior margin of seventh sternum emarginate ( Figs. 270 , 272 )............................................... 53

- Posterior margin of seventh sternum straight ( Fig. 273 )............................................. Hypatopa vitis

53. Posterior margin of seventh sternum laterally emarginate, forming a broad, truncate, median lobe ( Fig. 270 ).... Hypatopa rea

- Posterior margin of seventh sternum mesially and laterally emarginate, forming two rounded lobes ( Fig. 272 )... Hypatopa gena

54. Sex scales on membrane adjacent to Ostium bursae or on margin of seventh sternum ( Figs. 261–262 , 268 ).............. 55

- Sex scales absent ( Figs. 263–264 , 266–267 , 269, 271 )........................................................ 57

55. Sex scales on membrane adjacent to Ostium bursae ( Fig. 261 )..................................... Hypatopa umbra

- Sex scales on or near posterior margin of seventh sternum only ( Figs. 262 , 268 )................................... 56

56. Sex scales on posterolateral margin of seventh sternum; antrum present ( Fig. 262 )..................... Hypatopa hecate

- Sex scales along entire posterior margin of seventh segment; antrum absent ( Fig. 268 )................... Hypatopa rabio

57. Signum present ( Figs. 264 , 266 , 269, 271 )................................................................. 59

- Signum absent ( Fig. 263 , 267 ).......................................................................... 58

58. Eighth sternum subquadrate; ductus seminalis anterior to anterior margin of seventh sternum ( Fig. 267 )...... Hypatopa texo

- Eighth sternum longer than wide; ductus seminalis posterior to anterior margin of seventh sternum ( Fig. 263 )..................................................................................................... Hypatopa plebis

59. Eighth sternum quadrate ( Fig. 269 )............................................................. Hypatopa sais

- Eighth sternum longer than wide ( Figs. 264 , 266 , 271 )....................................................... 60

60. Antrum present ( Figs. 264 , 266 )......................................................................... 61

- Antrum absent ( Fig. 271 ).................................................................... Hypatopa hora

61. Antrum wider than long; anterior 1/4 of ductus bursae spinulate ( Fig. 266 )............................. Hypatopa leda

- Antrum longer than wide; anterior 1/2 of ductus bursae with two rows of internal platelets ( Fig. 264 )...... Hypatopa phoebe

Figures

FIGURES 135 – 140. Male genitalia of Blastobasinae. 135 – 136, Blastobasis beo (slide 3335). 137 – 138, Hypatopa nex (slide 2460). 139 – 140, Hypatopa cladis (slide 2345).

FIGURES 141 – 146. Male genitalia of Blastobasinae. 141 – 142, Hypatopa juno (slide 2037). 143 – 144, Hypatopa actes (slide 2640). 145 – 146, Hypatopa cotis (slide 2663).

FIGURES 147 – 152. Male genitalia of Blastobasinae. 147 – 148, Hypatopa pica (slide 2556). 149 – 150, Hypatopa hecate (slide 2029). 151 – 152, Hypatopa acus (slide 2004).

FIGURES 153 – 158. Male genitalia of Blastobasinae. 153 – 154, Hypatopa crux (slide 2041). 155 – 156, Hypatopa limae (slide 2691). 157 – 158, Hypatopa hera (slide 2667).

FIGURES 159 – 164. Male genitalia of Blastobasinae. 159 – 160, Hypatopa arxcis (slide 2664). 161 – 162, Hypatopa caedis (slide 2150). 163 – 164, Hypatopa plebis (slide 2210).

FIGURES 165 – 170. Male genitalia of Blastobasinae. 165 – 166, Hypatopa dolo (slide 2136). 167 – 168, Hypatopa cyane (slide 2398). 169 – 170, Hypatopa manus (slide 2566).

FIGURES 171 – 176. Male genitalia of Blastobasinae. 171 – 172, Hypatopa caepae (slide 2172). 173 – 174, Hypatopa cotytto (slide 2542). 175 – 176, Hypatopa lucina (slide 2401).

FIGURES 177 – 182. Male genitalia of Blastobasinae. 177 – 178, Hypatopa scobis (slide 2277). 179 – 180, Hypatopa agnae (slide 2220). 181 – 182, Hypatopa phoebe (slide 2544).

FIGURES 183 – 188. Male genitalia of Blastobasinae. 183 – 184, Hypatopa semela (slide 2586). 185 – 186, Hypatopa edax (slide 2325). 187 – 188, Hypatopa joniella (slide 83504).

FIGURES 189 – 194. Male genitalia of Blastobasinae. 189 – 190, Hypatopa rego (slide 2191). 191 – 192, Hypatopa styga (slide 2318). 193 – 194, Hypatopa texla (slide 2540).

FIGURES 195 – 200. Male genitalia of Blastobasinae. 195 – 196, Hypatopa verax (slide 2265). 197 – 198, Hypatopa tapadulcea Adamski, 1999 (slide 2069). 199 – 200, Hypatopa mora (slide 2567).

FIGURES 201 – 206. Male Genitalia of Blastobasinae. 201 – 202, Hypatopa nox (slide 2672). 203 – 204, Hypatopa dux (slide 2668). 205 – 206, Hypatopa erato (slide 2703).

FIGURES 207 – 212. Male genitalia of Blastobasinae. 207 – 208, Hypatopa fio (slide 2215). 209 – 210, Hypatopa io (slide 2372). 211 – 212, Hypatopa leda (slide 2284).

FIGURES 213 – 218. Male genitalia of Blastobasinae. 213 – 214, Hypatopa vox (slide 2020). 215 – 216, Hypatopa eos (slide 2723). 217 – 218, Hypatopa dicax (slide 2539).

FIGURES 219 – 224. Male genitalia of Blastobasinae. 219 – 220, Hypatopa ira (slide 2380). 221 – 222, Hypatopa umbra (slide 2168). 223 – 224, Hypatopa texo (slide 2263).

FIGURES 225 – 230. Male genitalia of Blastobasinae. 225 – 226, Hypatopa solea (slide 2601). 227 – 228, Hypatopa bilobata (slide 2595). 229 – 230, Hypatopa rabio (slide 2013).

FIGURES 231 – 236. Male genitalia of Blastobasinae. 231 – 232, Hypatopa rudis (slide 2449). 233 – 234, Hypatopa musa (slide 2392). 235 – 236, Pigritia sedis (slide 2722).

FIGURES 21 – 30. Wing venation of Blastobasinae. 21, Blastobasis vesta (slide 7051). 22, Blastobasis nivis (slide 7048). 23, Blastobasis phaedra (slide 7042). 24, Blastobasis aedes (slide 7050). 25, Blastobasis dicionis (slide 7039). 26, Hypatopa cladis (slide 7005). 27, Hypatopa juno (slide 7033). 28, Hypatopa pica (slide 7014). 29, Hypatopa hecate (slide 7015). 30, Hypatopa crux (slide 7035).

FIGURES 11 – 20. Wing venation of Blastobasinae. 11, Blastobasis erae (slide 7040). 12, Blastobasis iuanae (slide 7045). 13, Blastobasis graminea Adamski, 1999 (slide 83226). 14, Blastobasis neniae (slide 7055). 15, Blastobasis orithyia (slide 7053). 16, Blastobasis babae (slide 7049). 17, Blastobasis echus (slide 7036). 18, Blastobasis coffeaella (Busck, 1925) (USNM slide 83227). 19, Blastobasis rotullae (slide 7041). 20, Blastobasis deliciolarum (slide 7047).

FIGURES 263 – 265. Female genitalia of Blastobasinae. 263, Hypatopa plebis (Slide 4549). 264, Hypatopa phoebe (Slide 4525). 265, Hypatopa tapadulcea Adamski, 1999 (Taken from Adamski (2002).

FIGURES 269 – 271. Female genitalia of Blastobasinae. 269, Hypatopa sais (Slide 4500). 270, Hypatopa rea (Slide 4501). 271, Hypatopa hora (Slide 4503).

FIGURES 272 – 273. Female genitalia of Blastobasinae. 272, Hypatopa gena (Slide 2710). 273, Hypatopa vitis (Slide 4522).

FIGURES 260 – 262. Female genitalia of Blastobasinae. 260, Blastobasis dicionis (Slide 6007). 261, Hypatopa pica (Slide 4514). 262, Hypatopa hecate (Slide 4517).

FIGURES 266 – 268. Female genitalia of Blastobasinae. 266, Hypatopa leda (Slide 4512). 267, Hypatopa texo (Slide 4511). 268, Hypatopa rabio (Slide 4520).