Komiyandra lanyuana ( Hayashi, 1981 ), 2010

Santos-Silva, Antonio, Heffern, Daniel & Matsuda, Kiyoshi, 2010, Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae), Insecta Mundi 2010 (130), pp. 1-120 : 40-41

Komiyandra lanyuana ( Hayashi, 1981)

( Fig. 36, 37 , 96 , 158, 159 , 236 , 279 , 302, 308, 309 , 320 , 416-418 )

Parandra janus ; Kano 1938: 115, fig. 1.

Parandra lanyuana Hayashi, 1981: 27 ; Yu et al. 2002: 74, pl. 1, Fig. 7 a-7b; Mizuno and Shiyake 2004: 5, pl. 1, fig. 15 (types).

Birandra (Birandra) lanyuana ; Santos-Silva and Shute 2009: 32.

Description. Integument shining, dark-brown; parts of head, mandibles, scape, margins of pronotum, epipleura and elytral suture, and parts of legs blackish.

Male ( Fig. 416 ). Dorsal face of head, on gibbosities, coarsely and moderately abundantly punctate; longitudinal depression between gibbosities smooth; area behind gibbosities with transverse region smooth, interrupted in middle by punctures as coarse as on gibbosities, and with same kind of punctures between that region and occiput, but moderately sparse; area between gibbosities and ocular carina strongly depressed, coarsely punctate anteriorly and posteriorly; area behind eyes coarsely and sparsely punctate; ocular carina elevated, distinctly bifurcated in “Y” near posterior edge of eyes ( Fig. 416 ). Eyes narrow ( Fig. 96 ); posterior ocular edge ( Fig. 416 ) distinct. Central area of clypeus oblique close to front. Central projection of labrum ( Fig. 36 ) wide and slightly emarginate at apex. Submentum depressed from middle to anterior edge; punctation coarse, moderately sparse; pilosity moderately long, sparse; anterior edge elevated throughout. Mandibles approximately as long as head; teeth of inner margin ( Fig. 158 ) placed around middle. Antennae reaching basal fifth of the prothorax; ventral sensorial area of antennomeres III-XI not visible from side ( Fig. 236 ), and not divided by carina; dorsal sensorial area of antennomere XI moderately large, narrow and elliptical.

Lateral margins of prothorax subparallel at anterior two-thirds towards anterior angles. Pronotum finely, moderately abundantly punctate centrally, gradually coarser laterally, mainly close to anterior angles; lateral area microsculptured, more distinctly close to anterior angles; anterior edge slightly sinuous; anterior angles slightly projected forward; lateral angles not distinct; posterior angles distinct. Basal two-thirds of elytra coarsely, abundantly punctate, mainly laterally on basal third and across medial third; punctation of apical third finer, more abundant and concentrated; each elytron with two distinct carinae. Metasternum with punctures coarse and sparse laterally, gradually finer towards metasternal suture. Metafemur ( Fig. 417 ) moderately elongated. Dorsal face of metatibia slightly rounded at basal half, apical half flat; in dorsal view moderately wide. Metatarsomere V (without claws) slightly longer than I-III together ( Fig. 279 ); in dorsal view ( Fig. 302 ) not enlarged at basal half.

Male genitalia: median lobe long, forming a flattened tube, with two elongate struts at base, apex widely rounded and feebly incised at middle ( Fig. 309 ); parameres of median length, forming a ring, with a pair of pointed processes, separate near base in dorsal view, apical lobes long and wide, stout relatively narrow ( Fig. 308 ).

Female ( Fig. 418 ). Head moderately wide; dorsal face coarsely and abundantly punctate throughout. Central projection of labrum ( Fig. 37 ) narrow and slightly emarginate at apex. Mandible as in Fig. 159 . Pronotum with punctation as in male, but without distinct microsculptured lateral area; anterior angles more projected than in male; anterior edge sinuous. Antennae reaching basal fourth of prothorax. Metatibiae slightly sulcate on dorsal face of basal two-thirds.

Variability. Integument from brown to dark-brown. Male: central projection of labrum truncate at apex; submentum depressed throughout. Female: central projection of labrum truncate at apex.

Dimensions in mm (M / F). Total length (including mandibles), 16.9-20.5/15.1-19.5; prothorax: length, 4.0/3.5; anterior width, 4.8/4.2; posterior width, 3.9/3.9; humeral width, 4.8/4.6; elytral length, 9.7/9.8.

Geographical distribution. Taiwan (Lanyu Island).

Material examined. TAIWAN, Taitung: Lanyu Island , holotype M, VII.20.1976, H. Nara coll. ( OMNH) ; paratype F, same data of holotype ( HNCO) ; 2 M, 6 F, same data of holotype ( HNCO) ; M, F, VII.18.2001, W. L. Chen coll. ( HNCO); 1 M, 1 F, V.2002, [no collector indicated] ( NOCO). Specimen examined through photographs taken by Dr. Mei-Ling Chan ( NMNS): TAIWAN, Taitung: Lanyu Island (eclosed from pupa), M, V.1.2000, H. Y. Chu coll. ( NMNS) ; F, V.5.2000, H. Y. Chu coll. ( NMNS).

Types, type locality. Holotype M, preserved in the OMNH (ex-Collection Masao Hayashi), and paratype F, preserved in HNCO. Originally, both specimens belonged to HNCO. Mr. Hajime Nara (pers. comm.) stated that Dr. Hayashi retained the holotype in his personal collection.

Comments. Komiyandra lanyuana differs from K. formosana by the: head in female larger ( Fig. 417 ); apex of central projection of labrum of female truncate ( Fig. 37 ); metatarsomere V not enlarged on basal half ( Fig. 302 ) in both sexes. In K. formosana , head of female is proportionally smaller ( Fig. 415 ), apex of labrum is rounded ( Fig. 35 ), and metatarsomere V is distinctly enlarged on basal half ( Fig. 301 ) in both sexes. Differs from K. uenoi , mainly by metatibiae wider in dorsal view (distinctly narrower in K. uenoi ). See comments on K. shibatai .

Figures

Figure 1-44. Labrum. 1) Papuandra araucariae, male. 2) P. araucariae, female. 3) P. gressitti, holotype male. 4) P. weigeli, holotype male. 5) P. weigeli, paratype female. 6) P. queenslandensis, holotype female. 7) P. norfolkensis, holotype male. 8) P. norfolkensis, paratype female. 9) P. rothschildi, holotype male. 10) P. rothschildi, paratype female. 11) P. oberthueri, holotype female. 12) Caledonandra austrocaledonica, male. 13) C. austrocaledonica, female. 14) C. passandroides, male. 15) C. passandroides, female. 16) Storeyandra frenchi, male. 17) S. frenchi, female. 18) Malukandra heterostyla, holotype male. 19) M. jayawijayana, holotype male. 20) M. hornabrooki, holotype male. 21) M. hornabrooki, paratype female. 22) Melanesiandra striatifrons, male. 23) M. striatifrons, holotype female of Parandra vitiensis. 24) M. solomonensis, male. 25) M. solomonensis, female. 26) M. bougainvillensis, holotype male. 27) M.bougainvillensis, paratype female.28) M.birai, holotype male. 29) Stenandra kolbei, female. 30) Hawaiiandra puncticeps, male. 31) Komiyandra janus, female. 32) K. shibatai, male. 33) K. shibatai, female. 34) K. formosana, male. 35) K. formosana, female. 36) K. lanyuana, male. 37) K. lanyuana, female. 38) K. javana, holotype male. 39) K. javana, paratype female. 40) K. nayani, paratype male. 41) K. nayani, paratype female. 42) K. ohbayashii, holotype male. 43) K. ohbayashii, paratype female. 44) K. vivesi, holotype male.

Figure 90-104. Head, lateral view. 90) Papuandra queenslandensis, holotype female. 91) P. rothschildi, holotype male. 92) P. oberthueri, holotype female. 93) Stenandra kolbei, female. 94) Komiyandra shibatai, male. 95) K. formosana, male. 96) K. lanyuana, male. 97) K. javana, holotype male. 98) K. nayani, paratype male. 99) K. ohbayashii, holotype male. 100) K. luzonica, paratype male. 101) K. philippinensis, paratype male. 102) K. mindanao, holotype male. 103) K. mehli, paratype male. 104) K. vivesi, holotype male.

Figure 148-176. Mandibles, dorsal view. 148) Papuandra norfolkensis, holotype male (left mandible). 149) P. norfolkensis, holotype male (right mandible). 150) P. norfolkensis, paratype female. 151) P. rothschildi, holotype male. 152) P. rothschildi, paratype female. 153) Komiyandra janus, female. 154) K. shibatai, male. 155) K. shibatai, female. 156) K. formosana, male. 157) K. formosana, female. 158) K. lanyuana, male. 159) K. lanyuana female. 160) K. javana, holotype male. 161) K. javana, paratype female. 162) K. nayani, paratype male. 163) K. nayani, paratype female.164) K. ohbayashii, holotype male. 165) K. ohbayashii, paratype female. 166) K. luzonica, holotype male. 167) K. luzonica, paratype female. 168) K. philippinensis, holotype male. 169) K. philippinensis, paratype female. 170) K. mindanao, holotype male. 171) K. mindanao, paratype female. 172) K. mehli, holotype male. 173) K. mehli, paratype female. 174) K. vivesi, holotype male. 175) K. lombokia, holotype male. 176) K. lombokia, paratype female.

Figure 235-251. Antenna. 235) Komiyandra formosana, male. 236) K. lanyuana, male. 237) K. javana, holotype male. 238) K. nayani, paratype male. 239) K. ohbayashii, holotype male. 240) K. luzonica, paratype male. 241) K. philippinensis, paratype male. 242) K. mindanao, holotype male. 243) K. mehli, paratype male. 244) K. vivesi, holotype male. 245) K. lombokia, paratype male. 246) K. sulawesiana, paratype male. 247) K. irianjayana, holotype male. 248) K. menieri, paratype male. 249) K. sangihe, holotype male. 250) K. mindoro, holotype male. 251) K. niisatoi, holotype male.

Figure 277-299. Metatarsus. 277) Komiyandra shibatai, male. 278) K. formosana, male. 279) K. lanyuana, male. 280) K. javana, paratype male. 281) K. nayani, paratype male. 282) K. ohbayashii, holotype male. 283) K. luzonica, paratype male. 284) K. philippinensis, paratype male. 285) K. mindanao, holotype male. 286) K. mehli, paratype male. 287) K. vivesi, holotype male. 288) K. lombokia, paratype male. 289) K. sulawesiana, holotype male. 290) K. irianjayana, holotype male. 291) K. menieri, holotype male. 292) K. sangihe, holotype male. 293) K. mindoro, holotype male. 294) K. niisatoi, holotype male. 295) K. drumonti, holotype male. 296) K. cabigasi, paratype male. 297) K. koni, paratype male. 298) K. poggii, holotype male. 299) K. uenoi, paratype male.

Figure 300-314. 300-303) Metatarsomere V. 300) Komiyandra shibatai, male. 301) K. formosana, male. 302) K. lanyuana, male. 303) K. uenoi, holotype male. 304-311) Aedeagus. 304) K. shibatai, parameres. 305) K. shibatai, median lobe. 306) K. formosana, parameres. 307) K. formosana, median lobe. 308) K. lanyuana, parameres. 309) K. lanyuana, median lobe. 310) K. uenoi, paratype parameres. 311) K. uenoi, paratype median lobe. 312-314) Antennomere XI. 312) K. nayani, paratype male. 313) K. javana, holotype male. 314) Melanesiandra striatifrons, male.

Figure 317-322. Maps. 317) Malukandra hornabrooki. 318) Papuandra araucariae (circle), Komiyandra menieri (square), K. drumonti (diamond), Papuandra queenslandensis (star). 319) Caledonandra austrocaledonica. 320) Komiyandra formosana (circle), K. lanyuana (square). 321) Storeyandra frenchi. 322) Stenandra kolbei in Asia.

Figure 416-421. Habitus. 416) Komiyandra lanyuana, male, dorsal view. 417) K. lanyuana, male, ventral view. 418) K. lanyuana, female, dorsal view. 419) K. javana, holotype male, dorsal view. 420) K. javana, holotype male, ventral view. 421) K. javana, paratype female, dorsal view.

Figure 410-415. Habitus. 410) Komiyandra shibatai, male, dorsal view. 411) K. shibatai, male, ventral view. 412) K. shibatai, female, dorsal view. 413) K. formosana, male, dorsal view. 414) K. formosana, female, ventral view. 415) K. formosana, female, dorsal view.