Komiyandra menieri, Santos-Silva & Heffern & Matsuda, 2010

Santos-Silva, Antonio, Heffern, Daniel & Matsuda, Kiyoshi, 2010, Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae), Insecta Mundi 2010 (130), pp. 1-120 : 38-39

Komiyandra menieri sp. nov.

( Fig. 58, 59 , 108 , 180, 181 , 248 , 291 , 318 , 338 , 450-452 , 455 )

Parandra janus Bates, 1875: 47 ; Lameere 1902: 97 (Part).

Etymology. Dedicated to Dr. J. Menier (MNHN) for the loan of many specimens used in our study.

Type material. Holotype M, INDONESIA, New Guinea, Irian Jaya, West Papua: Hatam, D’Albertis coll. ( MCGD) . Paratypes (8 M, 7 F), as follows: INDONESIA, New Guinea, Irian Jaya, Papua: Biological Station of Pusppenssat (50 km S Nabire), M, I.13.1997, A. Weigel coll. ( AWCO) ; Andai, F (paralectotype of Parandra janus ), VIII.1872, D’Albertis coll. ( MCGD) ; M, (ex. Collection Argod), VIII.1872, L. M. D’Albertis coll. ( MNHN) ; F, (ex. Collection Bates), VIII.1872, L. M. D’Albertis coll. ( MNHN) ; Hatam, F, IX.1872, L. M. D’Albertis coll. ( MZSP) ; M, VI.1875, Beccari coll. ( MCGD). West Papua: Testega (1000-1300m), M, III.29-IV.2.1990, A. Riedel coll. ( MZSP). Maluku Islands, North Maluku: Ternate Island, Bruijn , F (ex- Collection R. Oberthür), [no date of collection], Bruijn coll. ( MNHN); (“ Acqui Conora ”) , 1 M, 1 F, 1874, Beccari coll. ( MCGD); 1 M, 1 F, XI.1874, Beccari coll. ( MCGD) ; 1 M, 1 F, (ex-Collection A. Argod), XI.1874, Beccari coll. ( MNHN). PAPUA NEW GUINEA, Morobe: Sattelberg, M (ex. Collection Fairmaire), [no date and collector indicated] ( MNHN) .

Description. Integument dark-brown; parts of head, of mandibles, margins of pronotum, elytral suture, and apical extreme of femora blackish.

Male ( Fig. 450 ). Dorsal surface of head, on gibbosities and between that and occiput, coarsely and abundantly punctate; area between gibbosities and ocular carina strongly depressed, smooth; area behind eyes coarsely punctate, with punctures sparser close to eyes; ocular carina elevated, clearly bifurcated in “Y” near posterior edge of eyes ( Fig. 452 ); area between bifurcation of ocular carina and eyes coarsely and abundantly punctate ( Fig. 452 ). Eyes narrow ( Fig. 108 ); posterior ocular edge ( Fig. 450 ) very distinct. Central area of clypeus almost vertical close to front. Central projection of labrum ( Fig. 58 ) wide and truncate at apex. Submentum depressed; punctation coarse and abundant; pilosity slightly long and sparse; anterior edge wide, elevated throughout. Mandibles approximately as long as head; teeth of inner margin ( Fig. 180 ) placed at middle. Ventral sensorial area of antennomeres III-XI not visible from side ( Fig. 248 ), and not divided by carina.

Pronotum finely punctate at central region, and gradually coarser and more abundant towards laterally, mainly close to anterior angles ( Fig. 453 ); anterior edge concave centrally; anterior angles slightly projected forward; lateral angles distinct, rounded; posterior angles distinct, obtuse. Elytra coarsely, abundantly punctate, mainly laterally on anterior two-thirds; each elytron with two barely visible carinae. Metasternum coarsely punctate laterally, with punctures more concentrated close to metepisterna. Metafemur ( Fig. 451 ) moderately short and enlarged. Dorsal face of metatibia flat throughout, except at base; in dorsal view moderately enlarged. Metatarsomere I not tumid, narrow in lateral view ( Fig. 291 ).

Female ( Fig. 455 ). Apex of central projection of labrum ( Fig. 59 ) truncate. Mandible as in Fig. 181 .

Variability. Dorsal face of head, on gibbosities, with punctures slightly fine and moderately sparse; punctation of dorsal face of head, between gibbosities and occiput, coarser and more abundant than on gibbosities; bifurcation of ocular carina slightly marked; pronotum finely punctate at central area; anterior angles of pronotum not projected forward. In specimens from Ternate Island, anterior angles of the pronotum, in both sexes, are more acute, and the lateral punctation of the pronotum, also in both sexes, is slightly sparser.

Dimensions in mm (M / F). Total length (including mandibles), 15.3-22.7/18.3-22.1; prothorax: length, 3.2-4.6/3.8-4.5; anterior width, 3.9-6.2/4.4-4.8; posterior width, 3.2-5.1/3.9-4.5; humeral width, 4.2-6.3/ 4.9-6.0; elytral length, 9.4-13.1/11.1-13.8.

Comments. Differs from K. philippinensis by the punctation of the head, close to the bifurcation of the ocular carina coarse and abundant, by the coarser lateral punctation of the pronotum, and by the punctation of the outer face of the ocular carina, close to the anterior ocular edge, with punctures moderately coarse and abundant. In K. philippinensis , the punctation of the area close to the bifurcation of the ocular carina, of the outer face of the ocular carina, and of the lateral of the pronotum is finer and sparser. See comments on K. irianjayana .

Figures

Figure 45-74. 45-73) Labrum. 45) Komiyandra luzonica, holotype male. 46) K. luzonica paratype female. 47) K. philippinensis, paratype male. 48) K. philippinensis, paratype female. 49) K. mindanao, holotype male. 50) K. mindanao, paratype female. 51) K. mehli, holotype male. 52) K. mehli, paratype female. 53) K. lombokia, holotype male. 54) K. lombokia, paratype female. 55) K. sulawesiana, holotype male. 56) K. sulawesiana, paratype female. 57) K. irianjayana, holotype male. 58) K. menieri, paratype male. 59) K. menieri, paralectotype female of P. janus. 60) K. sangihe, holotype male. 61) K. mindoro, holotype male. 62) K. niisatoi, holotype male. 63) K. niisatoi, paratype female. 64) K. drumonti, holotype male. 65) K. cabigasi, paratype male. 66) K. cabigasi, paratype female. 67) K. koni, paratype male. 68) K. koni, paratype female. 69) K. johkii, holotype male. 70) K. poggii, holotype male. 71) K. poggii, paratype female. 72) K. uenoi, paratype male. 73) K. uenoi, paratype female. 74) Hawaiiandra puncticeps, head dorsal view, female (CL, clypeus; LB, labrum; FR, frons; at, anterior tentorial pit; cl, clypeolabral suture; es, epistomal suture).

Figure 105-117. Head, lateral view. 105) K. lombokia, paratype male. 106) K. sulawesiana, holotype male. 107) K. irianjayana, holotype male. 108) K. menieri, paratype male. 109) K. sangihe, paratype male. 110) K. mindoro, holotype male. 111) K. niisatoi, paratype male. 112) K. drumonti, holotype male. 113) K. cabigasi, paratype male. 114) K. koni, paratype male. 115) K. johkii, holotype male. 116) K. poggii, holotype male. 117) K. uenoi, paratype male.

Figure 177-199. 177-195) Mandibles, dorsal view. 177) Komiyandra sulawesiana, holotype male. 178) K. sulawesiana, paratype female. 179) K. irianjayana, holotype male. 180) K. menieri, paratype male. 181) K. menieri, paratype female (paralectotype of Parandra janus). 182) K. sangihe, paratype male. 183) K. mindoro, holotype male. 184) K. niisatoi, holotype male. 185) K. niisatoi, paratype female. 186) K. drumonti, holotype male. 187) K. cabigasi, paratype male. 188) K. cabigasi, paratype female. 189) K. koni, paratype male. 190) K. koni, paratype female. 191) K. johkii, holotype male. 192) K. poggii, holotype male. 193) K. poggii, paratype female. 194) K. uenoi, paratype male. 195) K. uenoi, paratype female. 196-199) Maxillary palp. 196) Komiyandra javana, paratype male. 197) Caledonandra austrocaledonica, male. 198) C. passandroides, male. 199) Hawaiiandra puncticeps, male.

Figure 235-251. Antenna. 235) Komiyandra formosana, male. 236) K. lanyuana, male. 237) K. javana, holotype male. 238) K. nayani, paratype male. 239) K. ohbayashii, holotype male. 240) K. luzonica, paratype male. 241) K. philippinensis, paratype male. 242) K. mindanao, holotype male. 243) K. mehli, paratype male. 244) K. vivesi, holotype male. 245) K. lombokia, paratype male. 246) K. sulawesiana, paratype male. 247) K. irianjayana, holotype male. 248) K. menieri, paratype male. 249) K. sangihe, holotype male. 250) K. mindoro, holotype male. 251) K. niisatoi, holotype male.

Figure 277-299. Metatarsus. 277) Komiyandra shibatai, male. 278) K. formosana, male. 279) K. lanyuana, male. 280) K. javana, paratype male. 281) K. nayani, paratype male. 282) K. ohbayashii, holotype male. 283) K. luzonica, paratype male. 284) K. philippinensis, paratype male. 285) K. mindanao, holotype male. 286) K. mehli, paratype male. 287) K. vivesi, holotype male. 288) K. lombokia, paratype male. 289) K. sulawesiana, holotype male. 290) K. irianjayana, holotype male. 291) K. menieri, holotype male. 292) K. sangihe, holotype male. 293) K. mindoro, holotype male. 294) K. niisatoi, holotype male. 295) K. drumonti, holotype male. 296) K. cabigasi, paratype male. 297) K. koni, paratype male. 298) K. poggii, holotype male. 299) K. uenoi, paratype male.

Figure 317-322. Maps. 317) Malukandra hornabrooki. 318) Papuandra araucariae (circle), Komiyandra menieri (square), K. drumonti (diamond), Papuandra queenslandensis (star). 319) Caledonandra austrocaledonica. 320) Komiyandra formosana (circle), K. lanyuana (square). 321) Storeyandra frenchi. 322) Stenandra kolbei in Asia.

Figure 335-340. Maps. 335) Komiyandra mehli. 336) K. sulawesiana. 337) K. vivesi (circle), K. mindanao (square). 338) K. niisatoi (circle), K. janus (square), K. menieri (triangle). 339) K. sangihe. 340) K. lombokia.

Figure 447-453. Habitus. 447) Komiyandra sulawesiana, paratype female, dorsal view. 448) K. irianjayana, holotype male, dorsal view. 449) K. irianjayana, holotype male, ventral view. 450) K. menieri, paratype male, dorsal view. 451) K. menieri, paratype male, ventral view. 452) K. menieri, paratype male, punctation, head and pronotum. 453) K. philippinensis, paratype male, punctation, head and pronotum.

Figure 454-460. Habitus. 454) Komiyandra irianjayana, holotype male, punctation, head and pronotum. 455) K. menieri, paratype female, dorsal view (paralectotype of Parandra janus). 456) K. sangihe, holotype male, dorsal view. 457) K. sangihe, holotype male, ventral view. 458) K. mindoro, holotype male, dorsal view. 459) K. mindoro, holotype male, ventral view. 460) K. niisatoi, holotype male, dorsal view.