Komiyandra javana, Santos-Silva & Heffern & Matsuda, 2010

Santos-Silva, Antonio, Heffern, Daniel & Matsuda, Kiyoshi, 2010, Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae), Insecta Mundi 2010 (130), pp. 1-120 : 33-34

Komiyandra javana sp. nov.

( Fig. 38, 39 , 97 , 160, 161 , 196 , 237 , 280 , 313 , 328 , 419-420 )

Parandra janus (part); Lansberge 1884: 135; Lameere 1902: 97; Arigony 1984: 109.

Etymology. The name refers to the island of Java in Indonesia.

Type material. Holotype M, from INDONESIA, Java (Province of East Java): Malang, 1905, ex “de Moffarts” Collection ( IRSN) . Paratypes (11 M, 7 F), as follows: INDONESIA, Java: (locality unreadable), F (ex. Collection J. Waterstradt; ex. Collection Oberthür), 1904, [no collector indicated] ( MNHN) ; M, 28.XII.1934, ex. Tippmann Collection ( MZSP) ; M, [date not indicated], ex Donckier Collection ( IRSN) ; M, [date not indicated], ex “de Moffarts” Collection ( MZSP) ; 2 M, [date not indicated], ex Nonfried Collection ( IRSN) ; F, ex “de Moffarts” Collection ( IRSN) ; 1 M, 2 F, [date not indicated], V. M. Duchon coll. ( IRSN); M, [date not indicated] ( IRSN); Nongkedjadjar – Tengger (probably a place between Nongkedjadjar, East Java, and Tengger, West Java), M, 3.II.1934, ex Tippmann Collection ( USNM) ; M, 14.X.1934, ex Tippmann Collection ( USNM) ; F, 17.X.1934, ex Tippmann Collection ( USNM). East Java: Mount Kawi , 2 M, 2 F (ex. Collection Oberthür), 1898, J. B. Ledru coll. ( MNHN) .

Description. Integument shining, dark-brown; mandibles, parts of head, basal antennomeres, parts of legs, and edge of pro- and mesocoxal cavities darker.

Male ( Fig. 419 ). Head wide; dorsal surface, on gibbosities, with coarse punctures, not abundant or confluent; area close to eyes with large punctures, anastomosed; area between gibbosities and occiput with fine and sparse punctures; frontal gibbosities well marked, separated by a deep and wide furrow; area between gibbosities and ocular carina smooth and depressed; ocular carina elevated, not bifurcated in “Y” near posterior edge of eyes ( Fig. 419 ); area behind eyes smooth. Eyes narrow ( Fig. 97 ); posterior ocular edge ( Fig. 419 ) distinct. Central area of clypeus almost vertical close to front. Central projection of labrum ( Fig. 38 ) wide and truncate at apex. Submentum barely depressed, with moderately large, abundant punctures; margin close to mentum moderately wide and elevated; pilosity sparse, present throughout. Maxillary palp ( Fig. 196 ). Teeth of inner margin of mandibles ( Fig. 160 ) placed at middle. Ventral sensorial area of antennomeres III-XI not divided by carina ( Fig. 313 ) and not visible from the side ( Fig. 237 ).

Pronotum coarsely and abundantly punctate at sides (mainly at anterior half); center of disc finely, sparsely punctate; anterior edge ( Fig. 419 ) clearly sinuous; anterior angles feebly projected; lateral angles barely marked; posterior angles marked, subrounded (not distinct). Elytra somewhat coarsely, abundantly punctate (finer, sparser in the circum-scutellar area); each elytron with two low, clearly marked carinae. Metasternum moderately, coarsely punctate laterally. Metafemur ( Fig. 420 ) elongate. Dorsal face of tibiae flat at base, barely furrowed at apical half. Metatarsus (without claws) approximately as long as metatibia; metatarsomere V clearly longer than I-III together.

Female ( Fig. 421 ). Smooth area of head, between gibbosities and ocular carina, not depressed; punctures of dorsal surface as in male. Central area of clypeus almost oblique close to front. Central projection of labrum ( Fig. 39 ) narrow and truncate. Ocular carina not bifurcated. Ventral sensorial area of antennomeres III-XI and submentum as in male. Mandibles ( Fig. 161 ) with punctures and pilosity sparser than male. Pronotal punctures finer laterally than male; anterior angles clearly projected. Elytral punctures, elytral carinae, and dorsal face of the tibiae as in male, and with the same variations.

Variability. Integument from dark-brown to brown. Males: dorsal surface of head, on gibbosities, finely punctate with or without coarse punctures intermixed; dorsal face of head close to eyes with coarse punctures, not confluent; bifurcation in “Y” of ocular carina barely marked; area behind eyes with coarse and sparse punctures; central area of clypeus oblique close to front; submentum depressed; submentum not or barely elevated throughout close to mentum; center of disc of pronotum finely and abundantly punctate; anterior angles of pronotum projected forward; one or two elytral carina barely marked; basal third of tibiae rounded. Female: central area of clypeus clearly oblique close to front.

Dimensions in mm (M / F). Total length (including mandibles), 17.6-21.6/19.3-20.0; prothorax: length, 3.6-4.8/3.9-4.1; anterior width, 4.6-6.2/4.6-4.9; posterior width, 3.7-4.7/4.4; humeral width, 4.8-6.0/5.6- 5.7; elytral length, 10.5-12.6/12.2-12.4.

Comments. Males of K. javana differ from males K. philippinensis by the bifurcation of ocular carina barely marked or absent, by the dorsal face of the head close to the eyes (area of the bifurcation) coarsely punctate, usually confluent, by the posterior angles of the pronotum sub-rounded (not projected), and by antennomere III longer (1.1 times longer than the greatest width). In males of K. philippinensis , the ocular carina is clearly bifurcated, the punctures of the dorsal face of head close to eyes are fine and sparse, the posterior angles of pronotum are obtuse and projected, and antennomere III is shorter (length equal to the greatest width). Females of these species distinguished by antennomere III similar to males, and by the pattern of labrum: narrow and truncate in K. javana ; narrow and rounded in K. philippinensis .

Komiyandra javana differs from K. shibatai by the same differences of the males of K. philippinensis , by metatarsomere V clearly longer ( Fig. 280 ), and by the anterior half of the prothorax ( Fig. 419 ) with parallel sides. In K. shibatai , metatarsomere V ( Fig. 277 ) is shorter and the anterior half of prothorax ( Fig. 410 ) has divergent sides.

Differs from K. formosana by the central projection of labrum in female truncated at apex, and by metatarsomere V not enlarged at basal half. In K. formosana , the central projection of labrum in female is rounded at apex, and metatarsomere V is enlarged at basal half.

Differs from K. nayani and K. ohbayashii , mainly, by the apex of labrum of males wide and truncate. In K. nayani , the apex of labrum is narrow and subacute, and in K. ohbayashii it is narrow and rounded. From K. mehli , differs by metatarsomere V clearly longer, by the ocular carina of the male slightly or not bifurcated, by the head proportionally wider, by the antennae proportionally longer and antennomeres wider, and by the posterior angles of the pronotum sub-rounded. In K. mehli , metatarsomere V ( Fig. 291 ) is shorter, the antennae are proportionally shorter and the antennomeres narrower, and the posterior angles of the pronotum are obtuse and projected.

Finally, differs from K. luzonica and K. janus by the apex of labrum of the males wide and truncate ( Fig. 38 ), and by the sensorial area of the antennae not carinate. In K. luzonica and K. janus , the apex of labrum of the males ( Fig. 45 , 407 ) is narrow and subacute, and the sensorial area of the antennae is clearly carinate.

Lansberge (1884) wrote on Parandra janus : “C’est dans ma collection que se trouvent les exemplaires de Java. Ils ont été pris, le mâle sur le Mt. Gedeh, la femalle sur le Mt. Addjoeno, et ne diffèrent des individus des Molucques [Seram] et de Celebes que par la ponctuation des élytres un peu plus forte. Je ne puis done les considérer que comme des exemplaires d’une variété locale”. Lameere (1902) agreed with Lansberge (1884): “Il m’a été impossible, comme à van Lansberge, de trouver des différences spécifiques entre les exemplaires des Moluques, de la Nouvelle-Guinée et de Java ”. Following those authors, Parandra janus , has been recorded from Java (besides many other islands) by later authors. There is no doubt that the species mentioned by Lansberge (1884) and Lameere (1902) for Java, corresponds to K. javana (see differences between this species and K. janus ), and the species from Seram, cited by Lansberge (1884) corresponds to K. mehli .

Arigony (1984) examined two females from Java that also correspond to K. javana , although she figured a male that is very similar to the true Parandra janus . The description of Parandra janus made by Arigony (1984) corresponds, probably, to more than one species, and none of which is K. janus (= P. janus ).

Figures

Figure 1-44. Labrum. 1) Papuandra araucariae, male. 2) P. araucariae, female. 3) P. gressitti, holotype male. 4) P. weigeli, holotype male. 5) P. weigeli, paratype female. 6) P. queenslandensis, holotype female. 7) P. norfolkensis, holotype male. 8) P. norfolkensis, paratype female. 9) P. rothschildi, holotype male. 10) P. rothschildi, paratype female. 11) P. oberthueri, holotype female. 12) Caledonandra austrocaledonica, male. 13) C. austrocaledonica, female. 14) C. passandroides, male. 15) C. passandroides, female. 16) Storeyandra frenchi, male. 17) S. frenchi, female. 18) Malukandra heterostyla, holotype male. 19) M. jayawijayana, holotype male. 20) M. hornabrooki, holotype male. 21) M. hornabrooki, paratype female. 22) Melanesiandra striatifrons, male. 23) M. striatifrons, holotype female of Parandra vitiensis. 24) M. solomonensis, male. 25) M. solomonensis, female. 26) M. bougainvillensis, holotype male. 27) M.bougainvillensis, paratype female.28) M.birai, holotype male. 29) Stenandra kolbei, female. 30) Hawaiiandra puncticeps, male. 31) Komiyandra janus, female. 32) K. shibatai, male. 33) K. shibatai, female. 34) K. formosana, male. 35) K. formosana, female. 36) K. lanyuana, male. 37) K. lanyuana, female. 38) K. javana, holotype male. 39) K. javana, paratype female. 40) K. nayani, paratype male. 41) K. nayani, paratype female. 42) K. ohbayashii, holotype male. 43) K. ohbayashii, paratype female. 44) K. vivesi, holotype male.

Figure 90-104. Head, lateral view. 90) Papuandra queenslandensis, holotype female. 91) P. rothschildi, holotype male. 92) P. oberthueri, holotype female. 93) Stenandra kolbei, female. 94) Komiyandra shibatai, male. 95) K. formosana, male. 96) K. lanyuana, male. 97) K. javana, holotype male. 98) K. nayani, paratype male. 99) K. ohbayashii, holotype male. 100) K. luzonica, paratype male. 101) K. philippinensis, paratype male. 102) K. mindanao, holotype male. 103) K. mehli, paratype male. 104) K. vivesi, holotype male.

Figure 148-176. Mandibles, dorsal view. 148) Papuandra norfolkensis, holotype male (left mandible). 149) P. norfolkensis, holotype male (right mandible). 150) P. norfolkensis, paratype female. 151) P. rothschildi, holotype male. 152) P. rothschildi, paratype female. 153) Komiyandra janus, female. 154) K. shibatai, male. 155) K. shibatai, female. 156) K. formosana, male. 157) K. formosana, female. 158) K. lanyuana, male. 159) K. lanyuana female. 160) K. javana, holotype male. 161) K. javana, paratype female. 162) K. nayani, paratype male. 163) K. nayani, paratype female.164) K. ohbayashii, holotype male. 165) K. ohbayashii, paratype female. 166) K. luzonica, holotype male. 167) K. luzonica, paratype female. 168) K. philippinensis, holotype male. 169) K. philippinensis, paratype female. 170) K. mindanao, holotype male. 171) K. mindanao, paratype female. 172) K. mehli, holotype male. 173) K. mehli, paratype female. 174) K. vivesi, holotype male. 175) K. lombokia, holotype male. 176) K. lombokia, paratype female.

Figure 177-199. 177-195) Mandibles, dorsal view. 177) Komiyandra sulawesiana, holotype male. 178) K. sulawesiana, paratype female. 179) K. irianjayana, holotype male. 180) K. menieri, paratype male. 181) K. menieri, paratype female (paralectotype of Parandra janus). 182) K. sangihe, paratype male. 183) K. mindoro, holotype male. 184) K. niisatoi, holotype male. 185) K. niisatoi, paratype female. 186) K. drumonti, holotype male. 187) K. cabigasi, paratype male. 188) K. cabigasi, paratype female. 189) K. koni, paratype male. 190) K. koni, paratype female. 191) K. johkii, holotype male. 192) K. poggii, holotype male. 193) K. poggii, paratype female. 194) K. uenoi, paratype male. 195) K. uenoi, paratype female. 196-199) Maxillary palp. 196) Komiyandra javana, paratype male. 197) Caledonandra austrocaledonica, male. 198) C. passandroides, male. 199) Hawaiiandra puncticeps, male.

Figure 235-251. Antenna. 235) Komiyandra formosana, male. 236) K. lanyuana, male. 237) K. javana, holotype male. 238) K. nayani, paratype male. 239) K. ohbayashii, holotype male. 240) K. luzonica, paratype male. 241) K. philippinensis, paratype male. 242) K. mindanao, holotype male. 243) K. mehli, paratype male. 244) K. vivesi, holotype male. 245) K. lombokia, paratype male. 246) K. sulawesiana, paratype male. 247) K. irianjayana, holotype male. 248) K. menieri, paratype male. 249) K. sangihe, holotype male. 250) K. mindoro, holotype male. 251) K. niisatoi, holotype male.

Figure 277-299. Metatarsus. 277) Komiyandra shibatai, male. 278) K. formosana, male. 279) K. lanyuana, male. 280) K. javana, paratype male. 281) K. nayani, paratype male. 282) K. ohbayashii, holotype male. 283) K. luzonica, paratype male. 284) K. philippinensis, paratype male. 285) K. mindanao, holotype male. 286) K. mehli, paratype male. 287) K. vivesi, holotype male. 288) K. lombokia, paratype male. 289) K. sulawesiana, holotype male. 290) K. irianjayana, holotype male. 291) K. menieri, holotype male. 292) K. sangihe, holotype male. 293) K. mindoro, holotype male. 294) K. niisatoi, holotype male. 295) K. drumonti, holotype male. 296) K. cabigasi, paratype male. 297) K. koni, paratype male. 298) K. poggii, holotype male. 299) K. uenoi, paratype male.

Figure 300-314. 300-303) Metatarsomere V. 300) Komiyandra shibatai, male. 301) K. formosana, male. 302) K. lanyuana, male. 303) K. uenoi, holotype male. 304-311) Aedeagus. 304) K. shibatai, parameres. 305) K. shibatai, median lobe. 306) K. formosana, parameres. 307) K. formosana, median lobe. 308) K. lanyuana, parameres. 309) K. lanyuana, median lobe. 310) K. uenoi, paratype parameres. 311) K. uenoi, paratype median lobe. 312-314) Antennomere XI. 312) K. nayani, paratype male. 313) K. javana, holotype male. 314) Melanesiandra striatifrons, male.

Figure 323-328. Maps. 323) Komiyandra koni (circle), K. johkii (square), K. poggii (star). 324) K. nayani. 325) Caledonandra passandroides. 326) Komiyandra shibatai. 327) Melanesiandra solomonensis. 328) Komiyandra javana.

Figure 416-421. Habitus. 416) Komiyandra lanyuana, male, dorsal view. 417) K. lanyuana, male, ventral view. 418) K. lanyuana, female, dorsal view. 419) K. javana, holotype male, dorsal view. 420) K. javana, holotype male, ventral view. 421) K. javana, paratype female, dorsal view.

Figure 410-415. Habitus. 410) Komiyandra shibatai, male, dorsal view. 411) K. shibatai, male, ventral view. 412) K. shibatai, female, dorsal view. 413) K. formosana, male, dorsal view. 414) K. formosana, female, ventral view. 415) K. formosana, female, dorsal view.

Figure 45-74. 45-73) Labrum. 45) Komiyandra luzonica, holotype male. 46) K. luzonica paratype female. 47) K. philippinensis, paratype male. 48) K. philippinensis, paratype female. 49) K. mindanao, holotype male. 50) K. mindanao, paratype female. 51) K. mehli, holotype male. 52) K. mehli, paratype female. 53) K. lombokia, holotype male. 54) K. lombokia, paratype female. 55) K. sulawesiana, holotype male. 56) K. sulawesiana, paratype female. 57) K. irianjayana, holotype male. 58) K. menieri, paratype male. 59) K. menieri, paralectotype female of P. janus. 60) K. sangihe, holotype male. 61) K. mindoro, holotype male. 62) K. niisatoi, holotype male. 63) K. niisatoi, paratype female. 64) K. drumonti, holotype male. 65) K. cabigasi, paratype male. 66) K. cabigasi, paratype female. 67) K. koni, paratype male. 68) K. koni, paratype female. 69) K. johkii, holotype male. 70) K. poggii, holotype male. 71) K. poggii, paratype female. 72) K. uenoi, paratype male. 73) K. uenoi, paratype female. 74) Hawaiiandra puncticeps, head dorsal view, female (CL, clypeus; LB, labrum; FR, frons; at, anterior tentorial pit; cl, clypeolabral suture; es, epistomal suture).

Figure 403-409. Habitus. 403) Papuandra oberthueri, holotype female, dorsal view. 404) P. oberthueri, holotype female, ventral view. 405) Komiyandra janus, lectotype male, dorsal view. 406) K. janus, lectotype male, ventral view. 407) K. janus, lectotype male, head and prothorax. 408) K. janus, lectotype male, labels. 409) K. janus, female, dorsal view.