Caledonandra passandroides ( Thomson, 1867 )

Santos-Silva, Antonio, Heffern, Daniel & Matsuda, Kiyoshi, 2010, Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae), Insecta Mundi 2010 (130), pp. 1-120 : 49-50

Caledonandra passandroides ( Thomson, 1867)

( Fig. 14, 15 , 81 , 132, 133 , 198 , 213 , 221 , 264 , 325 , 373-375 )

Parandra passandroides Thomson, 1867: 107 , 116; Lameere 1902: 94, 104; Fauvel 1906: 40; Hayashi 1961b: 10; Webb 1994: 327 (note); Jenis 2008: 117.

Parandra Passandroides ; Thomson 1878: 4 (type).

Parandra (Parandra) passandroides ; Lameere 1913: 6 (cat.); 1919: 18; Arigony 1984: 114; Santos-Silva 2002: 32 (note).

Birandra (Birandra) passandroides ; Santos-Silva and Shute 2009: 32.

Description. Integument shining, dark-brown; dorsal and lateral contour of mandibles, and elytral suture blackish.

Male ( Fig. 373 ). Head wide, proportionally very large in relation to body size; dorsal surface nearly planar to sides of depression in “V”, without gibbosities or with gibbosities barely defined; punctation fine near clypeus, gradually coarser towards the occiput and areas near ocular carina, not confluent; area behind eyes with coarse punctures, deep, in part confluent; area between the depression in “V” and ocular carina, near clypeus, with circular depression, deep; ocular carina low, not bifurcated in “Y” near the posterior edge of the eyes ( Fig. 373 ). Eyes ( Fig. 81 ) moderately narrow. Labrum clearly oblique; central projection ( Fig. 14 ) strongly sculptured: two strong tubercles, projected forward; semi-circular projection, elevated, distinctly more prominent in relation to areas to sides and center (each tip of arc starts at the base of the tubercles, see again, Fig 14 ). Maxillary palp as in Fig. 198 . Submentum slightly elevated in central area, clearly separated from gula and part of gena, by semi-circular furrow, large moderately deep; punctation fine and sparse; edge close to mentum moderately narrow and elevated, preceded by large depression, deep throughout; pilosity very short and sparse. Mandibles as in Fig. 132 . Antennae ( Fig. 221 ) reaching or almost reaching basal fifth of prothorax.

Pronotum finely punctate laterally; center of disc with punctation very fine and sparse; anterior angles not projected forward; anterior margin concave. Elytra very finely punctate, mainly towards the suture; elytral carinae absent. Wings as in Fig. 213 . Metasternum with coarse, shallow punctures at sides. Metafemora as in Fig. 374 . Metatarsus (without claws) approximately as long as metatibia (including apical spines); metatarsomere V ( Fig. 264 ) as long as I-III together.

Female ( Fig. 375 ). Punctation of head like in males. Depression in “V” of the dorsal face of head shallower than in males. Labrum without tubercles and without semi-circular projection; apex ( Fig. 15 ) narrow and rounded. Submentum without transverse depression close to anterior edge. Mandibles as in Fig. 133 . Anterior angles of prothorax projected forward; anterior margin slightly sinuous (central area clearly concave).

Variability. Integument pale-brown to dark-brown; submentum completely blackish; legs brown. Males: depression semi-circular on dorsal face of head moderately shallow and barely defined; area behind eyes with punctures shallow; surface of submentum strongly furrowed transversely; area close to anterior margin of submentum with furrow narrow and very deep; submentum glabrous; center of pronotal disc smooth; metatarsus (without claws) longer than metatibia (length equals to 0.9 times). Female: depression in “V” of dorsal face of head like in males; submentum with transverse depression, shallow, close to anterior edge.

Dimensions in mm (M / F). Total length (including mandibles), 12.3-21.0/13.9-18.9; prothorax: length, 2.3-3.7/2.7-3.7; anterior width, 3.2-5.6/3.6-4.5; posterior width, 2.7-4.4/3.2-4.4; humeral width, 3.2-5.5/ 3.7-5.4; elytral length, 6.7-10.6/8.3-10.7.

Geographical distribution ( Fig. 325 ). New Caledonia (Grande Terre, Ouvéa, Lifou).

Material examined. (14 M, 10 F), as follows: new caledonia. F, [date not indicated], Deyr. coll. ( MCGD); F, [date not indicated] ( MCGD); 3 M, 4 F, [date not indicated] ( IRSN); 2 M, 1878, Deyr. coll. ( MCGD); M, F, XII.1-7.2006, Ivo Jenis coll. ( ZKCO). Grande Terre : Me Jejehari (750 m; 165 o 36’30”E, 21 o 28’45S), M, XI.5.1986, Brown coll. ( MEMU) ; Mount Panié (250 m), M, F, X.30.1986, Brown coll. ( MEMU) ; Ouril (60 miles of Nouméa), M, ex. Fauvel Collection, [date not indicated] ( MZSP) ; Sarraméa , M, F, XII.27.2006- I.10.2007, Ivo Jenis coll. ( ZKCO) ; Tonghoué , M, F, [date not indicated] ( IRSN) ; Table Unio Road (21 o 34’S, 165 o 46’E; 600 m), 2 M, 14.XI.2000, Bouchard, Burwell, Monteith coll. ( QMCO). Lifou: M, [no date indicated] ( IRSN) .

Type, type locality. Holotype M, from New Caledonia, deposited at MNHN.

Comments. One of the authors (K. Matsuda) photographed three “types” in the collection of the MNHN: 1 lectotype male; 1 paralectotype male; and 1 paralectotype female. The designations of lectotype and paralectotype were never published. Therefore, the labels fixed on each of the specimens have no value. Besides, Thomson (1867) wrote: “Long 17 mill.; lat. 5 mill.” That indicates that Thomson (1867) had just one specimen, because when he had more than one, he indicated the measurements, for example, in Parandra sayi described in the same work: “Long. 11-17 mill.; lat. 3 1/2-5 1/2 mill.”. Thomson (1867) also did not know the female of P. passandroides , which, by itself, immediately excludes the “ paralectotype ” female. Thomson (1867) also stated at the end of his description that he had one specimen given to him by Fauvel. Besides, Thomson (1867) wrote: “prothorax sublaevis, antice quase rectus” [English translation: “prothorax almost smooth, anteriorly almost straight”]. That description of the anterior margin of the prothorax fits better to the male that has the label of “ Lectotype ” than the other. We believe that the holotype is the specimen that has the label of “Type” (characteristic of the types of the J. Thomson Collection) and “ Lectotype ”. The other two specimens that have labels of “ paralectotype ” should not be considered as types.

Figures

Figure 1-44. Labrum. 1) Papuandra araucariae, male. 2) P. araucariae, female. 3) P. gressitti, holotype male. 4) P. weigeli, holotype male. 5) P. weigeli, paratype female. 6) P. queenslandensis, holotype female. 7) P. norfolkensis, holotype male. 8) P. norfolkensis, paratype female. 9) P. rothschildi, holotype male. 10) P. rothschildi, paratype female. 11) P. oberthueri, holotype female. 12) Caledonandra austrocaledonica, male. 13) C. austrocaledonica, female. 14) C. passandroides, male. 15) C. passandroides, female. 16) Storeyandra frenchi, male. 17) S. frenchi, female. 18) Malukandra heterostyla, holotype male. 19) M. jayawijayana, holotype male. 20) M. hornabrooki, holotype male. 21) M. hornabrooki, paratype female. 22) Melanesiandra striatifrons, male. 23) M. striatifrons, holotype female of Parandra vitiensis. 24) M. solomonensis, male. 25) M. solomonensis, female. 26) M. bougainvillensis, holotype male. 27) M.bougainvillensis, paratype female.28) M.birai, holotype male. 29) Stenandra kolbei, female. 30) Hawaiiandra puncticeps, male. 31) Komiyandra janus, female. 32) K. shibatai, male. 33) K. shibatai, female. 34) K. formosana, male. 35) K. formosana, female. 36) K. lanyuana, male. 37) K. lanyuana, female. 38) K. javana, holotype male. 39) K. javana, paratype female. 40) K. nayani, paratype male. 41) K. nayani, paratype female. 42) K. ohbayashii, holotype male. 43) K. ohbayashii, paratype female. 44) K. vivesi, holotype male.

Figure 75-89. Head, lateral view, male. 75) Hawaiiandra puncticeps. 76) Storeyandra frenchi. 77) Malukandra heterostyla. 78) M. jayawijayana, holotype. 79) M. hornabrooki, holotype. 80) Caledonandra austrocaledonica. 81) C. passandroides. 82) Melanesiandra striatifrons. 83) M. solomonensis, paratype. 84) M. bougainvillensis, holotype. 85) M. birai, holotype. 86) Papuandra araucariae. 87) P. gressitti, holotype. 88) P. weigeli, holotype. 89) P. norfolkensis, holotype.

Figure 118-147. Mandibles, dorsal view. 118) Birandra (Yvesandra) angulicollis, male. 119) B. (Y.) angulicollis, female. 120) Acutandra araucana, male. 121) Parandra (Parandra) glabra, female. 122) Stenandra kolbei, female. 123) Hawaiiandra puncticeps, male. 124) Storeyandra frenchi, male. 125) S. frenchi, female. 126) Malukandra heterostyla, holotype male. 127) M. jayawijayana, holotype male. 128) M. hornabrooki, holotype male. 129) M. hornabrooki, paratype female. 130) Caledonandra austrocaledonica, male. 131) C. austrocaledonica, female. 132) C. passandroides, male. 133) C. passandroides, female. 134) Melanesiandra striatifrons, male. 135) M. striatifrons, female. 136) M. solomonensis, male. 137) M. solomonensis, female. 138) M. bougainvillensis, holotype male. 139) M. bougainvillensis, paratype female. 140) M. birai, holotype male. 141) Papuandra araucariae, paratype male. 142) P. araucariae, female. 143) P. gressitti, holotype male. 144) P. weigeli, holotype male. 145) P. weigeli, paratype female. 146) P. queenslandensis, holotype male. 147) P. oberthueri, holotype female.

Figure 177-199. 177-195) Mandibles, dorsal view. 177) Komiyandra sulawesiana, holotype male. 178) K. sulawesiana, paratype female. 179) K. irianjayana, holotype male. 180) K. menieri, paratype male. 181) K. menieri, paratype female (paralectotype of Parandra janus). 182) K. sangihe, paratype male. 183) K. mindoro, holotype male. 184) K. niisatoi, holotype male. 185) K. niisatoi, paratype female. 186) K. drumonti, holotype male. 187) K. cabigasi, paratype male. 188) K. cabigasi, paratype female. 189) K. koni, paratype male. 190) K. koni, paratype female. 191) K. johkii, holotype male. 192) K. poggii, holotype male. 193) K. poggii, paratype female. 194) K. uenoi, paratype male. 195) K. uenoi, paratype female. 196-199) Maxillary palp. 196) Komiyandra javana, paratype male. 197) Caledonandra austrocaledonica, male. 198) C. passandroides, male. 199) Hawaiiandra puncticeps, male.

Figure 210-217. 210-214) Wings. 210) Papuandra araucariae, male. 211) Komiyandra nayani, paratype male. 212) Birandra (Birandra) silvaini, male. 213) Caledonandra passandroides, male. 214) Melanesiandra striatifrons, male. 215-217) Antenna. 215) Stenandra kolbei, female. 216) Hawaiiandra puncticeps, male. 217) Storeyandra frenchi, male.

Figure 218-234. Antenna. 218) Malukandra jayawijayana, holotype male. 219) M. hornabrooki, paratype female. 220) Caledonandra austrocaledonica, male. 221) C. passandroides, male. 222) Melanesiandra striatifrons, male. 223) M. solomonensis, male. 224) M. bougainvillensis, holotype male. 225) M. birai, holotype male.226) Papuandra araucariae, male. 227) P. gressitti, holotype male. 228) P. weigeli, holotype male. 229) P. queenslandensis, holotype female. 230) P. norfolkensis, holotype male. 231) P. rothschildi, paratype male. 232) P. oberthueri, holotype female. 233) Komiyandra janus, female. 234) K. shibatai, male.

Figure 252-276. 252-257) Antenna. 252) Komiyandra drumonti, holotype male. 253) K. cabigasi, paratype male. 254) K. johkii, holotype male. 255) K. poggii, holotype male. 256) K. koni, paratype male. 257) K. uenoi, paratype male. 258-276) Metatarsus. 258) Stenandra kolbei, female. 259) Hawaiiandra puncticeps, male. 260) Storeyandra frenchi, male. 261) Malukandra jayawijayana, holotype male. 262) M. hornabrooki, holotype male. 263) Caledonandra austrocaledonica, male. 264) C. passandroides, male. 265) Melanesiandra striatifrons, male. 266) M. solomonensis, male. 267) M. bougainvillensis, holotype male. 268) M. birai, holotype male. 269) Papuandra araucariae, male. 270) P. gressitti, holotype male. 271) P. weigeli, holotype male. 272) P. queenslandensis, holotype female. 273) P. norfolkensis, holotype male. 274) P. rothschildi, holotype male. 275) oberthueri, holotype female. 276) Komiyandra janus, female.

Figure 323-328. Maps. 323) Komiyandra koni (circle), K. johkii (square), K. poggii (star). 324) K. nayani. 325) Caledonandra passandroides. 326) Komiyandra shibatai. 327) Melanesiandra solomonensis. 328) Komiyandra javana.

Figure 373-378. Habitus. 373) Caledonandra passandroides, male, dorsal view. 374) C. passandroides, male, ventral view. 375) C. passandroides, female, dorsal view. 376) Melanesiandra striatifrons, male, dorsal view. 377) M. striatifrons, male, ventral view. 378) M. striatifrons, holotype female of Parandra vitiensis.