Komiyandra, Santos-Silva & Heffern & Matsuda, 2010

Santos-Silva, Antonio, Heffern, Daniel & Matsuda, Kiyoshi, 2010, Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae), Insecta Mundi 2010 (130), pp. 1-120 : 9-12

Key to the species of Komiyandra

1. Sensorial area of antennomeres III-XI divided by carina, or with carina in some of those antennomeres ........................................................................................................................... 2

— Sensorial area of antennomeres III-XI not divided by carina ...................................................... 9

2(1). Elytral punctation fine or moderately fine ( Fig. 428, 430 , 445 , 447 ) .......................................... 3

— Elytral punctation coarse ( Fig. 405, 409 , 410, 412 ) .................................................................... 4

3(2). Posterior angles of pronotum rounded ( Fig. 445 , 447 ) in both sexes. Indonesia [Sulawesi, Java (?)] ........................................................................................................ K. sulawesiana , sp. nov.

— Posterior angles of pronotum distinct and obtuse in both sexes ( Fig. 428, 430 ). Philippines (Luzon, Mindoro, Mindanao) ............................................................................... K. luzonica , sp. nov.

4(2). Metatarsomere V shorter than half the length of metatibiae. Japan (Ryukyu Islands: Amami-Ôshima and Okinawa Is.) .................................................................... K. shibatai (Hayashi)

— Metatarsomere V longer than half the length of metatibiae ....................................................... 5

5(4). Lateral margins of pronotum ( Fig. 441 , 466 , 468 ) distinctly expanded ...................................... 6

— Lateral margins of pronotum ( Fig. 422, 424 ) not distinctly expanded ....................................... 7

6(5). Dorsal face of metatibiae narrowed at apical half; metatarsomere V (without claws) as long as I-III together. Philippines (Mindanao) ........................................................... K. vivesi , sp. nov.

— Dorsal face of metatibiae wide at apical half; metatarsomere V (without claws) longer than I-III together. Philippines (Mindanao) .......................................................... K. cabigasi , sp. nov.

7(5). Elytral punctation notably abundant and close ( Fig. 422, 424 ), the general appearance of elytral surface somewhat rugose; pronotal punctation in female distinctly abundant laterally, close to anterior angles. Malaysia (Sabah) ............................................................ K. nayani , sp. nov.

— Elytral punctation abundant but not notably close ( Fig. 425, 427 ); the general appearance of elytral surface not rugose; pronotal punctation in female sparse or moderately sparse laterally, close to anterior angles ............................................................................................................. 8

8(7). Ocular carina in male not bifurcated in “Y” near posterior edge of eyes; lateral margins of anterior two-thirds of pronotum in female ( Fig. 427 ) convergent towards head. Philippines (Mindanao) ........................................................................................................... K. ohbayashii , sp. nov.

— Ocular carina in male bifurcated in “Y” near posterior edge of eyes; lateral margins of anterior two-thirds of pronotum in female ( Fig. 409 ) parallel. Indonesia (Sulawesi). K. janus (Bates)

9(1). Metatarsomere I notably tumid, wide in lateral view ( Fig. 290 ). Indonesia (Irian Jaya) .............. ........................................................................................................ K. irianjayana , sp. nov.

— Metatarsomere I not tumid and narrow in lateral view ( Fig. 288 ) ........................................... 10

10(9). Elytral punctation fine. Indonesia (Lombok, Sumbawa Island) ............. K. lombokia , sp. nov.

— Elytral punctation coarse .......................................................................................................... 11

11(10). Central projection of labrum ( Fig. 40, 41 ) narrow, acute or sub-acute at apex in both sexes. Malaysia (Sabah) ...................................................................................... K. nayani , sp. nov.

— Central projection of labrum of male ( Fig. 34 , 61 ) wide, truncate or slightly rounded; in female, narrow, truncate ( Fig. 73 ) or acute ( Fig. 33 ) (in this last case, species not present in Borneo) ................................................................................................................................................ 12

12(11). Elytra distinctly flat. Body clearly compressed dorsoventrally ( Fig. 465 ). Papua New Guinea.... .............................................................................................................. K. drumonti , sp. nov.

— Elytra convex. Body not compressed dorsoventrally ( Fig. 433 ) ................................................. 13

13(12). Metatarsomere V not notably narrow at basal half ( Fig. 301 ). Taiwan (excluding Lanyu Island) ........................................................................................ K. formosana (Miwa and Mitono)

— Metatarsomere V notably narrow at basal half ( Fig. 300 ) ........................................................ 14

14(13). Mandibles subtriangular ( Fig. 155 , 193 ); length, at most, one-third longer than the greater width. Female .................................................................................................................................... 28

— Mandibles sub-falciform ( Fig. 154 , 184 ); length, at least, equal to 1.5 times the greater width. Male ........................................................................................................................................ 15

15(14). Teeth of inner margin of mandible in male ( Fig. 183 ) placed after middle (occupying only apical third of inner margin) Female unknown. Philippines (Mindoro) .......... K. mindoro , sp. nov.

— Teeth of inner margin of mandible in male ( Fig. 189 ) placed in middle (occupying great part of inner margin) ......................................................................................................................... 16

16(15). Head narrower than pronotum, elongate behind eyes ( Fig. 438 ) .............................................. 17

— Head as wide as pronotum, not elongate behind eyes ( Fig. 431 ) ............................................... 21

17(16). Metepisternae and urosternites coarsely and abundantly punctate ......................................... 18

— Metepisternae and urosternites finely and somewhat sparsely punctate ................................. 20

18(17). Central area of labrum distinctly tumid and with tuberculiform process frontally; central projection of labrum not notably projected at apex, and strongly lowered. Indonesia (Sumatra) ............... .................................................................................................................... K. poggii , sp. nov.

— Central area of labrum not tumid and without tuberculiform process ..................................... 19

19(18). Dorsal face of metatibiae flat; latero-outer face of metatibiae longitudinally sulcate. Borneo ( Malaysia and Indonesia) ............................................................................................... K. koni , sp. nov.

— Dorsal face of tibiae rounded; latero-outer face of metatibiae not sulcate. Borneo ( Malaysia) ....... .................................................................................................................... K. johkii , sp. nov.

20(17). Integument very dark brown (blackish). Female unknown. Indonesia (Sulawesi) ....................... ................................................................................................................. K. niisatoi , sp. nov.

— Integument dark-brown (not blackish). Indonesia (Seram) .......................... K. mehli , sp. nov.

21(16). Ocular carina not bifurcated in “Y” near posterior edge of eyes, or with bifurcation vaguely indicated ................................................................................................................................................ 22

— Ocular carina distinctly bifurcated in “Y” near posterior edge of eyes ..................................... 24

22(21). Anterior edge of pronotum ( Fig. 419 ) sinuous (emargination very distinct at middle). Indonesia (Java) ........................................................................................................ K. javana , sp. nov.

— Anterior edge of pronotum ( Fig. 456 ) not sinuous (wide and slightly sinuous at middle) ........ 23

23(22). Greatest width of metafemur less than three times the length. Male. Indonesia (Sulawesi) ....... ................................................................................................................ K. sangihe , sp. nov.

— Greatest width of metafemur slightly more than three times the length. Philippines (Mindanao) ............................................................................................................ K. mindanao , sp. nov.

24(21). Lateral margins of anterior third of pronotum divergent towards head. Japan (Ryukyu Islands: Amami-Ôshima Islands and Okinawa Islands) .................................. K. shibatai (Hayashi)

— Lateral margins of anterior third of pronotum parallel or subparallel ..................................... 25

25(24). Metatibiae narrow in dorsal view (lateral not notably visible at medial region). Japan (Ryukyu Islands: Ishigaki-jima Island, Iriomote-jima Island) ................................... K. uenoi , sp. nov.

— Metatibiae moderately wide in dorsal view (lateral notably visible at medial region) .............. 25

26(25). Area behind bifurcation of ocular carina with punctures abundant and at least partially anastomosed, or networked. Indonesia (Irian Jaya, Ternate Island)] ..... K. menieri , sp. nov.

— Area behind bifurcation of ocular carina with punctures sparse .............................................. 27

27(26). Metatarsomere V approximately as long as half the length of metatibia. Philippines (Negros Island, Leyte Island, Sibuyan Island, Luzon Island, Mindoro Island) ........................................ ..................................................................................................... K. philippinensis , sp. nov.

— Metatarsomere V distinctly shorter than half the length of metatibia. Taiwan (Lanyu Island) .. .......................................................................................................... K. lanyuana (Hayashi)

28(14). Central projection of labrum with apex narrow and acute (triangular) ( Fig. 33 , 48 ) ............... 29

— Central projection of labrum truncate or distinctly rounded ( Fig. 39 , 68 ) ............................... 30

29(28). Head ( Fig. 412 ) distinctly narrower than pronotum. Japan (Ryukyu Islands: Amami-Ôshima Islands and Okinawa Islands) ............................................................. K. shibatai (Hayashi)

— Head ( Fig. 434 ) wide, extending nearly to lateral edge of pronotum. Philippines (Negros Island, Leyte Island, Sibuyan Island, Luzon Island, Mindoro Island) .... K. philippinensis , sp. nov.

30(28). Metatarsomere V longer than I-III together .............................................................................. 31

— Metatarsomere V shorter or as long as I-III together ............................................................... 34

31(30). Head ( Fig. 421 ) wide (lateral distance between apices of eyes, greater than 2.5 times the length of mandible in dorsal view) Indonesia (Java) ............................................... K. javana , sp. nov.

— Head ( Fig. 472 ) narrow (lateral distance between apices of eyes, at most, 2.5 times the length of mandible in dorsal view) ......................................................................................................... 32

32(31). Circum-scutellar punctation not different of that placed laterally on basal third of elytra. Borneo ( Malaysia and Indonesia) ............................................................................... K. koni , sp. nov.

— Circum-scutellar punctation distinctly different of that placed laterally on basal third of elytra ................................................................................................................................................ 33

33(32). Metepisterna with punctures coarse and shallow. Indonesia (Irian Jaya, Ternate Island), Papua New Guinea............................................................................................. K. menieri , sp. nov.

— Metepisterna with punctures slightly coarse and well defined. Indonesia (Seram) ....................... .................................................................................................................... K. mehli , sp. nov.

34(30). Head elongate behind eyes ( Fig. 437 ) ........................................................................................ 35

— Head not elongate behind eyes ( Fig. 480 ) .................................................................................. 36

35(34). Metepisternae abundantly punctate throughout, but the punctures are closer at basal third. Indonesia (Sumatra) ................................................................................... K. poggii , sp. nov.

— Metepisterna not abundantly punctate. Philippines (Mindanao) .......... K. mindanao , sp. nov.

36(34). Metatibiae, in dorsal view, notably narrow at apical half. Japan (Ryukyu Islands: Ishigaki-jima Island, Iriomote-jima Island) ....................................................................... K. uenoi , sp. nov.

— Metatibiae, in dorsal view, not notably narrow at apical half. Taiwan (Lanyu Island) ................ .......................................................................................................... K. lanyuana (Hayashi)

Figures

Figure 428-434. Habitus. 428) Komiyandra luzonica, paratype male, dorsal view. 429) K. luzonica, paratype male, ventral view. 430) K. luzonica, paratype female, dorsal view. 431) K. philippinensis, paratype male, dorsal view. 432) K. philippinensis, paratype male, ventral view. 433) K. philippinensis, paratype male, lateral view. 434) K. philippinensis, paratype female, dorsal view.

Figure 441-446. Habitus. 441) Komiyandra vivesi, holotype male, dorsal view. 442) K. lombokia, holotype male, dorsal view. 443) K. lombokia, holotype male, ventral view. 444) K. lombokia, paratype female, dorsal view. 445) K. sulawesiana, holotype male, dorsal view. 446) K. sulawesiana, holotype male, ventral view.

Figure 447-453. Habitus. 447) Komiyandra sulawesiana, paratype female, dorsal view. 448) K. irianjayana, holotype male, dorsal view. 449) K. irianjayana, holotype male, ventral view. 450) K. menieri, paratype male, dorsal view. 451) K. menieri, paratype male, ventral view. 452) K. menieri, paratype male, punctation, head and pronotum. 453) K. philippinensis, paratype male, punctation, head and pronotum.

Figure 403-409. Habitus. 403) Papuandra oberthueri, holotype female, dorsal view. 404) P. oberthueri, holotype female, ventral view. 405) Komiyandra janus, lectotype male, dorsal view. 406) K. janus, lectotype male, ventral view. 407) K. janus, lectotype male, head and prothorax. 408) K. janus, lectotype male, labels. 409) K. janus, female, dorsal view.

Figure 410-415. Habitus. 410) Komiyandra shibatai, male, dorsal view. 411) K. shibatai, male, ventral view. 412) K. shibatai, female, dorsal view. 413) K. formosana, male, dorsal view. 414) K. formosana, female, ventral view. 415) K. formosana, female, dorsal view.

Figure 461-466. Habitus. 461) Komiyandra niisatoi, holotype male, ventral view. 462) K. niisatoi, paratype female, dorsal view. 463) K. drumonti, holotype male, dorsal view. 464) K. drumonti, holotype male, ventral view. 465) K. drumonti, holotype male, lateral view. 466) K. cabigasi, holotype male, dorsal view.

Figure 467-472. Habitus. 467) Komiyandra cabigasi, holotype male, ventral view. 468) K. cabigasi, paratype female, ventral view. 469) K. koni, paratype male, dorsal view. 470) K. koni, paratype male, ventral view. 471) K. koni, paratype male, lateral view. 472) K. koni, paratype female, dorsal view. 473) K. johkii, holotype male, dorsal view.

Figure 422-427. Habitus. 422) Komiyandra nayani, holotype male, dorsal view. 423) K. nayani, holotype male, ventral view. 424) K. nayani, paratype female, dorsal view. 425) K. ohbayashii, holotype male, dorsal view. 426) K. ohbayashii, holotype male, ventral view. 427) K. ohbayashii, paratype female, dorsal view.

Figure 277-299. Metatarsus. 277) Komiyandra shibatai, male. 278) K. formosana, male. 279) K. lanyuana, male. 280) K. javana, paratype male. 281) K. nayani, paratype male. 282) K. ohbayashii, holotype male. 283) K. luzonica, paratype male. 284) K. philippinensis, paratype male. 285) K. mindanao, holotype male. 286) K. mehli, paratype male. 287) K. vivesi, holotype male. 288) K. lombokia, paratype male. 289) K. sulawesiana, holotype male. 290) K. irianjayana, holotype male. 291) K. menieri, holotype male. 292) K. sangihe, holotype male. 293) K. mindoro, holotype male. 294) K. niisatoi, holotype male. 295) K. drumonti, holotype male. 296) K. cabigasi, paratype male. 297) K. koni, paratype male. 298) K. poggii, holotype male. 299) K. uenoi, paratype male.

Figure 1-44. Labrum. 1) Papuandra araucariae, male. 2) P. araucariae, female. 3) P. gressitti, holotype male. 4) P. weigeli, holotype male. 5) P. weigeli, paratype female. 6) P. queenslandensis, holotype female. 7) P. norfolkensis, holotype male. 8) P. norfolkensis, paratype female. 9) P. rothschildi, holotype male. 10) P. rothschildi, paratype female. 11) P. oberthueri, holotype female. 12) Caledonandra austrocaledonica, male. 13) C. austrocaledonica, female. 14) C. passandroides, male. 15) C. passandroides, female. 16) Storeyandra frenchi, male. 17) S. frenchi, female. 18) Malukandra heterostyla, holotype male. 19) M. jayawijayana, holotype male. 20) M. hornabrooki, holotype male. 21) M. hornabrooki, paratype female. 22) Melanesiandra striatifrons, male. 23) M. striatifrons, holotype female of Parandra vitiensis. 24) M. solomonensis, male. 25) M. solomonensis, female. 26) M. bougainvillensis, holotype male. 27) M.bougainvillensis, paratype female.28) M.birai, holotype male. 29) Stenandra kolbei, female. 30) Hawaiiandra puncticeps, male. 31) Komiyandra janus, female. 32) K. shibatai, male. 33) K. shibatai, female. 34) K. formosana, male. 35) K. formosana, female. 36) K. lanyuana, male. 37) K. lanyuana, female. 38) K. javana, holotype male. 39) K. javana, paratype female. 40) K. nayani, paratype male. 41) K. nayani, paratype female. 42) K. ohbayashii, holotype male. 43) K. ohbayashii, paratype female. 44) K. vivesi, holotype male.

Figure 45-74. 45-73) Labrum. 45) Komiyandra luzonica, holotype male. 46) K. luzonica paratype female. 47) K. philippinensis, paratype male. 48) K. philippinensis, paratype female. 49) K. mindanao, holotype male. 50) K. mindanao, paratype female. 51) K. mehli, holotype male. 52) K. mehli, paratype female. 53) K. lombokia, holotype male. 54) K. lombokia, paratype female. 55) K. sulawesiana, holotype male. 56) K. sulawesiana, paratype female. 57) K. irianjayana, holotype male. 58) K. menieri, paratype male. 59) K. menieri, paralectotype female of P. janus. 60) K. sangihe, holotype male. 61) K. mindoro, holotype male. 62) K. niisatoi, holotype male. 63) K. niisatoi, paratype female. 64) K. drumonti, holotype male. 65) K. cabigasi, paratype male. 66) K. cabigasi, paratype female. 67) K. koni, paratype male. 68) K. koni, paratype female. 69) K. johkii, holotype male. 70) K. poggii, holotype male. 71) K. poggii, paratype female. 72) K. uenoi, paratype male. 73) K. uenoi, paratype female. 74) Hawaiiandra puncticeps, head dorsal view, female (CL, clypeus; LB, labrum; FR, frons; at, anterior tentorial pit; cl, clypeolabral suture; es, epistomal suture).

Figure 300-314. 300-303) Metatarsomere V. 300) Komiyandra shibatai, male. 301) K. formosana, male. 302) K. lanyuana, male. 303) K. uenoi, holotype male. 304-311) Aedeagus. 304) K. shibatai, parameres. 305) K. shibatai, median lobe. 306) K. formosana, parameres. 307) K. formosana, median lobe. 308) K. lanyuana, parameres. 309) K. lanyuana, median lobe. 310) K. uenoi, paratype parameres. 311) K. uenoi, paratype median lobe. 312-314) Antennomere XI. 312) K. nayani, paratype male. 313) K. javana, holotype male. 314) Melanesiandra striatifrons, male.

Figure 148-176. Mandibles, dorsal view. 148) Papuandra norfolkensis, holotype male (left mandible). 149) P. norfolkensis, holotype male (right mandible). 150) P. norfolkensis, paratype female. 151) P. rothschildi, holotype male. 152) P. rothschildi, paratype female. 153) Komiyandra janus, female. 154) K. shibatai, male. 155) K. shibatai, female. 156) K. formosana, male. 157) K. formosana, female. 158) K. lanyuana, male. 159) K. lanyuana female. 160) K. javana, holotype male. 161) K. javana, paratype female. 162) K. nayani, paratype male. 163) K. nayani, paratype female.164) K. ohbayashii, holotype male. 165) K. ohbayashii, paratype female. 166) K. luzonica, holotype male. 167) K. luzonica, paratype female. 168) K. philippinensis, holotype male. 169) K. philippinensis, paratype female. 170) K. mindanao, holotype male. 171) K. mindanao, paratype female. 172) K. mehli, holotype male. 173) K. mehli, paratype female. 174) K. vivesi, holotype male. 175) K. lombokia, holotype male. 176) K. lombokia, paratype female.

Figure 177-199. 177-195) Mandibles, dorsal view. 177) Komiyandra sulawesiana, holotype male. 178) K. sulawesiana, paratype female. 179) K. irianjayana, holotype male. 180) K. menieri, paratype male. 181) K. menieri, paratype female (paralectotype of Parandra janus). 182) K. sangihe, paratype male. 183) K. mindoro, holotype male. 184) K. niisatoi, holotype male. 185) K. niisatoi, paratype female. 186) K. drumonti, holotype male. 187) K. cabigasi, paratype male. 188) K. cabigasi, paratype female. 189) K. koni, paratype male. 190) K. koni, paratype female. 191) K. johkii, holotype male. 192) K. poggii, holotype male. 193) K. poggii, paratype female. 194) K. uenoi, paratype male. 195) K. uenoi, paratype female. 196-199) Maxillary palp. 196) Komiyandra javana, paratype male. 197) Caledonandra austrocaledonica, male. 198) C. passandroides, male. 199) Hawaiiandra puncticeps, male.

Figure 435-440. Habitus. 435) Komiyandra mindanao, holotype male, dorsal view. 436) K. mindanao, holotype male, ventral view. 437) K. mindanao, paratype female. 438) K. mehli, holotype male, dorsal view. 439) K. mehli, paratype male, ventral view. 440) K. mehli, paratype female, dorsal view.

Figure 416-421. Habitus. 416) Komiyandra lanyuana, male, dorsal view. 417) K. lanyuana, male, ventral view. 418) K. lanyuana, female, dorsal view. 419) K. javana, holotype male, dorsal view. 420) K. javana, holotype male, ventral view. 421) K. javana, paratype female, dorsal view.

Figure 454-460. Habitus. 454) Komiyandra irianjayana, holotype male, punctation, head and pronotum. 455) K. menieri, paratype female, dorsal view (paralectotype of Parandra janus). 456) K. sangihe, holotype male, dorsal view. 457) K. sangihe, holotype male, ventral view. 458) K. mindoro, holotype male, dorsal view. 459) K. mindoro, holotype male, ventral view. 460) K. niisatoi, holotype male, dorsal view.

Figure 474-480. Habitus. 474) Komiyandra johkii, holotype male, ventral view. 475) K. poggii, holotype male, dorsal view. 476) K. poggii, holotype male, ventral view. 477) K. poggii, paratype female, dorsal view. 478) K. uenoi, paratype male, dorsal view. 479) K. uenoi, paratype male, ventral view. 480) K. uenoi, paratype female, dorsal view.