Komiyandra luzonica, Santos-Silva & Heffern & Matsuda, 2010

Santos-Silva, Antonio, Heffern, Daniel & Matsuda, Kiyoshi, 2010, Revision of Hawaiian, Australasian, Oriental, and Japanese Parandrinae (Coleoptera, Cerambycidae), Insecta Mundi 2010 (130), pp. 1-120 : 13-14

Komiyandra luzonica sp. nov.

( Fig. 45, 46 , 100 , 166, 167 , 240 , 283 , 333 , 428-430 )

Etymology. The name refers to the island of Luzon in the Philippines.

Type material. Holotype M, Philippines, Cordillera Administrative Region , luzon, Benguet: Baguio, [no date indicated], G. G. Haslam col. ( USNM) . Paratypes (22 M, 17 F), as follows: PHILIPPINES, Cordillera Administrative Region, Luzon, Mountain Province: Mount Puguis (1800 – 1900 m), 2 M, VII.17.1985, M. Sakai coll. ( EELE) ; Mount Polis, F, V.2006, D. Mohagan coll. ( KMCT). Ifugao: Banaue, F, VI.1987, [no collector indicated] ( HNCO). Benguet: M, [no date indicated], Sanio Tomas coll. ( USNM) ; Baguio, 2 M, F, [no date indicated], G. G. Haslam coll. ( USNM) ; M, [no date indicated], G. G. Haslam col. ( MZSP). Cagayan Valley Region, Nueva Viscaya : 2 F, VI.2004, local coll. ( EVCO) ; M, [no date indicated], I. Luwawig coll. ( EVCO); Santa Fe, 10 M, 10 F, VI.2006, local. coll. ( DHCO) ; M, F, VI.2006, local. coll. ( MSZP). Mimaropa Region, Mindoro , Oriental Mindoro: Mount Halcon, M, VI.2007, [no collector indicated] ( MZSP) ; M, V.2001, [no collector indicated] ( ZKCO) ; M, VI.2007, [no collector indicated] ( ZKCO). Northern Mindanao Region, Mindanao , Bukidnon: Mount Kalatungan, M, F, IX.2007, ( ZKCO) .

Description. Integument shining, dark-brown; ventral surface and tarsus chestnut.

Male ( Fig. 428 ). Head wide; dorsal surface, on gibbosities, with barely fine punctures, moderately abundant, not confluent; area between gibbosities and occiput with same kind of punctures, but sparser; area close to eyes with barely large punctures, confluent; frontal gibbosities well marked, separated by deep and wide furrow; area between gibbosities and ocular carina depressed, and with some punctures; ocular carina elevated, not bifurcated in “Y” near posterior edge of eyes ( Fig. 428 ); area behind eyes with punctures slightly coarse, sparse. Eyes narrow ( Fig. 100 ); posterior ocular edge ( Fig. 428 ) distinct. Central area of clypeus vertical close to front. Central projection of labrum ( Fig. 45 ) narrow and subacute at apex. Submentum barely depressed, with large and abundant punctures; margin close to mentum wide and barely elevated at middle (more elevated at sides); pilosity very sparse, present throughout. Teeth of inner margin of mandibles ( Fig. 166 ) placed after middle. Ventral sensorial area of antennomeres III-XI divided by carina, visible from side ( Fig. 240 ).

Pronotum coarsely, abundantly punctate at latero-basal half; latero-apical half with tiny, abundant callosities; center of disc finely, sparsely punctate; anterior edge ( Fig. 428 ) clearly sinuous; anterior angles not projected forward; lateral angles barely marked; posterior angles marked, obtuse and barely distinct. Elytra somewhat finely, moderately, sparsely punctate, mainly towards suture; each elytron with two carinae barely visible. Metasternum with some moderately coarse, shallow punctures at sides. Metafemur ( Fig. 429 ) elongated. Dorsal face of tibiae rounded; in dorsal view distinctly wide. Metatarsus (without claws) approximately as long as metatibia; metatarsomere V barely longer than I-III together.

Female ( Fig. 430 ). Latero-apical half of pronotum with abundant punctures, and moderately rugose. Labrum ( Fig. 46 ). Mandible ( Fig. 167 ).

Variability. Males: area between gibbosities and ocular carina with fine, sparse punctures; area close to eyes with punctures partially confluent; area between gibbosities and ocular carina smooth; ocular carina bifurcated in “Y” near posterior edge of eyes; central projection of labrum narrow and rounded at apex; punctures of submentum moderately abundant; margin of submentum, close to mentum, wide and elevated throughout; pilosity sparse; submentum depressed at sides; anterior margin of pronotum barely sinuous; anterior angles of pronotum barely projected forward; elytral punctures fine and sparse, mainly towards suture; each elytron with a single carina barely visible. Female: Latero-apical half of pronotum finely, sparsely punctate, and not rugose.

Dimensions in mm (M / F). Total length (including mandibles), 16.0-23.1/17.2-18.0; prothorax: length, 3.3-4.8/3.5-3.7; anterior width, 3.9-6.5/4.2-4.3; posterior width, 3.3-5.1/3.9-4.0; humeral width, 4.3-6.4/ 5.0-5.1; elytral length, 9.5-12.7/11.0-11.6.

Comments. Differs from males of K. shibatai ( Hayashi, 1963) , K. lanyuana ( Hayashi, 1981) , K. formosana ( Miwa and Mitono, 1939) , K. philippinensis and K. mehli , mainly, by the central projection of labrum narrow and rounded or subacute at apex, and by the elytral punctures fine or nearly fine, distinctly sparse. In males of the species mentioned above, the apex of the central projection of labrum is wide and truncate, and the elytral punctures are distinctly coarser and close in both sexes.

Differs from K. nayani , K. ohbayashii and K. janus , by the presence of tiny callosities at latero-apical half of the pronotum, and by the elytral punctures clearly finer. In the three species, the pronotum is only punctate at latero-apical half, and their elytral punctures are clearly coarser and more abundant than elytral punctures of K. luzonica .

Figures

Figure 45-74. 45-73) Labrum. 45) Komiyandra luzonica, holotype male. 46) K. luzonica paratype female. 47) K. philippinensis, paratype male. 48) K. philippinensis, paratype female. 49) K. mindanao, holotype male. 50) K. mindanao, paratype female. 51) K. mehli, holotype male. 52) K. mehli, paratype female. 53) K. lombokia, holotype male. 54) K. lombokia, paratype female. 55) K. sulawesiana, holotype male. 56) K. sulawesiana, paratype female. 57) K. irianjayana, holotype male. 58) K. menieri, paratype male. 59) K. menieri, paralectotype female of P. janus. 60) K. sangihe, holotype male. 61) K. mindoro, holotype male. 62) K. niisatoi, holotype male. 63) K. niisatoi, paratype female. 64) K. drumonti, holotype male. 65) K. cabigasi, paratype male. 66) K. cabigasi, paratype female. 67) K. koni, paratype male. 68) K. koni, paratype female. 69) K. johkii, holotype male. 70) K. poggii, holotype male. 71) K. poggii, paratype female. 72) K. uenoi, paratype male. 73) K. uenoi, paratype female. 74) Hawaiiandra puncticeps, head dorsal view, female (CL, clypeus; LB, labrum; FR, frons; at, anterior tentorial pit; cl, clypeolabral suture; es, epistomal suture).

Figure 90-104. Head, lateral view. 90) Papuandra queenslandensis, holotype female. 91) P. rothschildi, holotype male. 92) P. oberthueri, holotype female. 93) Stenandra kolbei, female. 94) Komiyandra shibatai, male. 95) K. formosana, male. 96) K. lanyuana, male. 97) K. javana, holotype male. 98) K. nayani, paratype male. 99) K. ohbayashii, holotype male. 100) K. luzonica, paratype male. 101) K. philippinensis, paratype male. 102) K. mindanao, holotype male. 103) K. mehli, paratype male. 104) K. vivesi, holotype male.

Figure 148-176. Mandibles, dorsal view. 148) Papuandra norfolkensis, holotype male (left mandible). 149) P. norfolkensis, holotype male (right mandible). 150) P. norfolkensis, paratype female. 151) P. rothschildi, holotype male. 152) P. rothschildi, paratype female. 153) Komiyandra janus, female. 154) K. shibatai, male. 155) K. shibatai, female. 156) K. formosana, male. 157) K. formosana, female. 158) K. lanyuana, male. 159) K. lanyuana female. 160) K. javana, holotype male. 161) K. javana, paratype female. 162) K. nayani, paratype male. 163) K. nayani, paratype female.164) K. ohbayashii, holotype male. 165) K. ohbayashii, paratype female. 166) K. luzonica, holotype male. 167) K. luzonica, paratype female. 168) K. philippinensis, holotype male. 169) K. philippinensis, paratype female. 170) K. mindanao, holotype male. 171) K. mindanao, paratype female. 172) K. mehli, holotype male. 173) K. mehli, paratype female. 174) K. vivesi, holotype male. 175) K. lombokia, holotype male. 176) K. lombokia, paratype female.

Figure 235-251. Antenna. 235) Komiyandra formosana, male. 236) K. lanyuana, male. 237) K. javana, holotype male. 238) K. nayani, paratype male. 239) K. ohbayashii, holotype male. 240) K. luzonica, paratype male. 241) K. philippinensis, paratype male. 242) K. mindanao, holotype male. 243) K. mehli, paratype male. 244) K. vivesi, holotype male. 245) K. lombokia, paratype male. 246) K. sulawesiana, paratype male. 247) K. irianjayana, holotype male. 248) K. menieri, paratype male. 249) K. sangihe, holotype male. 250) K. mindoro, holotype male. 251) K. niisatoi, holotype male.

Figure 277-299. Metatarsus. 277) Komiyandra shibatai, male. 278) K. formosana, male. 279) K. lanyuana, male. 280) K. javana, paratype male. 281) K. nayani, paratype male. 282) K. ohbayashii, holotype male. 283) K. luzonica, paratype male. 284) K. philippinensis, paratype male. 285) K. mindanao, holotype male. 286) K. mehli, paratype male. 287) K. vivesi, holotype male. 288) K. lombokia, paratype male. 289) K. sulawesiana, holotype male. 290) K. irianjayana, holotype male. 291) K. menieri, holotype male. 292) K. sangihe, holotype male. 293) K. mindoro, holotype male. 294) K. niisatoi, holotype male. 295) K. drumonti, holotype male. 296) K. cabigasi, paratype male. 297) K. koni, paratype male. 298) K. poggii, holotype male. 299) K. uenoi, paratype male.

Figure 329-334. Maps. 329) Melanesiandra bougainvillensis. 330) Malukandra jayawijayana (circle), Papuandra weigeli (square), Komiyandra irianjayana (star). 331) Papuandra gressitti (circle), P. rothschildi (square). 332) Komiyandra ohbayashii. 333) K. luzonica. 334) K. philippinensis.

Figure 428-434. Habitus. 428) Komiyandra luzonica, paratype male, dorsal view. 429) K. luzonica, paratype male, ventral view. 430) K. luzonica, paratype female, dorsal view. 431) K. philippinensis, paratype male, dorsal view. 432) K. philippinensis, paratype male, ventral view. 433) K. philippinensis, paratype male, lateral view. 434) K. philippinensis, paratype female, dorsal view.