Manota occulta, Hippa, Heikki & Papp, László, 2007

Hippa, Heikki & Papp, László, 2007, The genus Manota Williston (Diptera: Mycetophilidae) in Thailand, with the description of seven new species, Zootaxa 1528, pp. 41-60 : 56-58

Manota occulta sp. n.

( Figs 9 A, B)

Body dark brown, length ca. 2.7 mm, wing length ca. 2.10 mm (defective).

Male. Head. Flagellomere 4 as long as broad, its trichia 0.04 mm. Postocular setae (8) medium-long, black. Thorax. Anepisternum setose. Anterior basalare setose. Setae of preepisternum 2 not observed. Laterotergite setose. Setae of episternum 3 not observed. Mid and hind trochanters, basal ¼ of mid and hind femora dark, ventral surface of fore femur fumose, tarsi brown. Wing (right wing lost, left wing damaged). Membrane dorsally without setae at posterior margin, veins brown. Sc distally of h non-setose. Ratio of visible sections of veins M1 and M2 48/ca. 105. A1 indicated by a few setae.

Hypopygium ( Fig. 9 A, B). Tergite 9 subtriangular, posteriorly scarcely extending to half length of gonocoxa, laterally sharply demarcated, anteriorly deeply incised, apically not distinctly separated from more posterior membraneous structures, the setae stronger than on gonocoxae. Mesial margin of gonocoxa sigmoid, with a small apicomesial lobe. Parastylar lobe not exposed but covered by the apicomesial lobe of gonocoxa, weakly sclerotized, with ca. 10 posteriorly directed setae. Paraapodemal lobe indistinct, not visible in ventral view. Dorsal mesial margin of gonocoxa subapically with a rounded lobe with a few setae and more apically with a larger posteriorly directed lobe, which is setose apically and along the mesial margin; apically on gonocoxa at the base of the gonostylus with a few setae arising from large basal bodies. One juxtagonostylar seta present: a stout, possibly flattened, megaseta arising from a basal body which is about a third of the length of the megaseta. Gonostylus about twice as long as broad, broadest at middle, apically slightly bilobate on ventral side, setosity moderately long, rather unmodified. Tegmen triangular, with weak lateral shoulders. Hypoproct rather large, posteriorly extending as far as gonocoxa, with ca 20 scattered ventral setae. Cerci medially fused.

Female unknown.

Discussion. Manota occulta is similar to M. secreta . It is distinguished by the juxtagonostylar megaseta which is simple, not flattened, by having the ventral mesial margin of gonocoxa sigmoid, not almost straight, by having the lobe at the ventral apicomesial corner of gonocoxa more pronounced, by having the more anterior of the lobes on the dorsal mesial margin of the gonocoxa more pronounced, by having the apical margin of the gonostylus oblique, not transversely emarginated, and by having the gonostylus apicolaterally bilobed. Furthermore, the parastylar lobe has ca. 10 setae instead of ca. 5. Both species are similar to M. roslii , M. yongi and M. pollex . They differ from M. roslii e.g. by having a prominent lobe at the dorsal apicomesial corner of the gonocoxa. This lobe is setose on its margin and to a variable extent on the dorsal surface. In M. roslii the homologous place is only slightly produced lobe-like and bears setae on the ventral surface. M. occulta and M. secreta differ from M. pollex and M. yongi e.g. by lacking a long and strong seta at the dorsal mesial margin of the gonocoxa, situated on a lobe which is apparently homologous with the more anterior of the two lobes in these species.

Types. Holotype. Male, Thailand: Doi Pui, over a forest trail, No. 10, Oct 31, 2004, leg. L. Papp & M. Földvári (gen. prep. Bud 22).

Figures

FIGURE 9. Manota occulta sp. n. (holotype): hypopygium in dorsal (A) and in ventral (B) view. Scale 0.1 mm.