Mallinella melanognatha ( van Hasselt, 1882 ) Dankittipakul & Jocqué & Singtripop, 2012

Dankittipakul, Pakawin, Jocqué, Rudy & Singtripop, Tippawan, 2012, Systematics and biogeography of the spider genus Mallinella Strand, 1906, with descriptions of new species and new genera from Southeast Asia (Araneae, Zodariidae) 3369, Zootaxa 3369 (1), pp. 1-327 : 217-218

Mallinella melanognatha ( van Hasselt, 1882) comb. nov.

(Figs 986–987, 1008, 1016–1017, 1020–1031)

Storena melanognatha van Hasselt, 1882: 34 ; plate 2, fig. 6, pl 5, figs 1–2, description of ♀. Kulczyṅski, 1911b: 485, plate 21, fig. 33, description of ♀. Kritscher, 1957: 266; figs 22–24, description of ♂ (misidentification = male of M. calilungae ). Thorell, 1897: 192, description of ♀ (misidentification)

Material examined. Holotype: ♀, INDONESIA, Sumatra, RMHN 5115, examined. Original label: / Sum: Exp [= Sumatra Expedition]; Storena ; melanognatha ; ♀ v. H [= van Hasselt]/ from middle Sumatra. Old material. INDONESIA, Sumatra, South Sumatra Province: 1♀ ( NMMV, examined). New material. INDONESIA, Sumatra, Lampung Province: 1♂, Mt. Barisan Selatan NP, road Krui – Liwa, 2 km southwest of Kubuprahu ,

480–540 m, disturbed evergreen rainforest and secondary forest, 4 March 2000, leg. P.J. Schwendinger ( MHNG, Sum–00/25). Singkep Islands : 1♂, 1♀, Batu Ampar Waterfall, ca. 10 km northwest of Dabo (00º29'30.9"S, 104º28'31.4"E) , 80 m, rainforest along stream, 17–18 June 2001, leg. P.J. Schwendinger ( MHNG, SIM –01/04). Lingga Islands : 1♀, at the foot of Mt. Daik, ca. 5 km NW of Daik (00º12'35.9S, 104º36'58.8"E) , 50 m, rainforest along stream, 12 June 2001, leg. P.J. Schwendinger ( MHNG, SIM –01/02) .

Taxonomic remarks. The holotype of S. melanognatha and a female mentioned by Kulczyṅski (1911) were collected on Sumatra. A conspecific male previously designated and described as ‘ Storena melanognatha ’ by Kritscher (1957) was found on Luzon, the Philippines. This is also the case in a male identified by Simon (deposited in the collection of the MNHN, AR 1559). The males erroneously recognized as ‘ S. melanognatha ’ by the latter two taxonomists belong to M. calilungae (NMMV) and M. simoni Dankittipakul et al., 2011 ( MNHN). The male of M. melanognatha comb. nov. is described here for the first time.

Diagnosis. Females of M. melanognatha comb. nov. can be recognized by the broad epigynal plate and by the spherical apical part of the spermathecae. Males are distinguished by the triangular embolic base with concave anterior margin, and by the trapezoidal flange on the embolus.

Redescription. Female (holotype). Measurements see van Hasselt (1882).

Coloration (Figs 1020–1021). Carapace orange-brown. Chelicerae yellowish brown. Sternum orange-brown. Legs pale yellow. Opisthosoma discolored; dorsum whitish, with faint pigmentation of pale brown. Dorsal pattern: anteriorly provided with faint marking of reniform patches, followed by two oval spots and a transverse chevron.

Genitalia. (Fig. 1022). Epigynal plate trapezoidal, anterior margin slightly wider than posterior one. Lateral borders sharply pointed, projecting inwards.

Female (new material, MHNG). Total length 7.20; prosoma 3.80 long, 3.00 wide. Eye sizes and interdistances: AME 0.28, ALE 0.22, PME 0.20, PLE 0.22, AME–AME 0.26, AME–ALE 0.36, PME–PME 0.24, PME–PLE 0.42; MOQ: 0.58 long, 0.46 anterior width, 0.40 posterior width. Leg measurements: I 14.34 (3.44, 4.50, 3.60, 2.78), II 11.88 (3.00, 3.74, 2.98, 2.16), III 11.96 (3.14, 3.60, 3.20, 2.00), IV 16.54 (3.98, 4.86, 5.16, 2.48).

Coloration (Fig. 987). Carapace dark chestnut-brown. Chelicerae dark brown. Sternum brown. Legs yellowish, femora distally brown. Dorsum of opisthosoma dark brown. Dorsal pattern: first and second pairs fused, forming reniform patches; third to fifth pairs represented by paired oval spots.

Genitalia (Figs 1008, 1016–1017, 1028–1031). Epigynal plate simple, both sides parallel. Internal genitalia with moderately long insemination ducts distinctly separated from spermathecae. Spermathecae divided into two parts: basal part cylindrical, apical part spherical.

Male (here designated). Total length 7.50; prosoma 3.90 long, 3.00 wide. Eye sizes and interdistances: AME 0.30, ALE 0.22, PME 0.20, PLE 0.22, AME–AME 0.28, AME–ALE 0.40, PME–PME 0.26, PME–PLE 0.50; MOQ: 0.60 long, 0.52 anterior width, 0.46 posterior width. Leg measurements: I 14.10 (3.38, 4.42, 3.54, 2.74), II 11.68 (2.96, 3.68, 2.92, 2.12), III 11.76 (3.10, 3.54, 3.16, 2.00), IV 16.24 (3.92, 4.78, 5.06, 2.46).

Coloration (Fig. 986). Carapace dark chestnut-brown. Chelicerae dark brown. Sternum brown. Legs yellowish. Dorsum of opisthosoma dark brown. Dorsal pattern: first and second pairs fused, forming reniform patches; third to fifth pairs represented by paired oval spots.

Palp (Figs 1024–1027). RTA digitiform, apex blunt. TA rostrated; apical ridge quadrangular, occupying half length of TA; apico-prolateral process slender, sharply pointed, directed posteromesad; baso-retrolateral ridge with blunt basal tooth. Embolic base triangular, anterior margin strongly excavated; membranous area narrowed, almost indistinct. Embolus broad at base, gradually tapered towards apex, medially with short trapezoidal flange.

Distribution. Sumatra and islands to the east.