Lathrobium (s.str.) geminum Kraatz, 1857

Ryvkin, Alexandr B., 2011, On new and poorly known Lathrobium (s. str.) species from Siberia and the Russian Far East (Insecta: Coleoptera: Staphylinidae: Paederinae), Baltic Journal of Coleopterology 11 (2), pp. 135-170 : 147-150

Lathrobium (s.str.) geminum Kraatz, 1857

( Figs. 5–7 )

Lathrobium geminum Kraatz, 1857: 673

Lat hrobium geminum ; Fauvel, 1873: 343 Lathrobium (Lathrobium) geminum ; Czwalina, 1888, Deutsche Entomologische Zeitschrift, 32(2): 344

Lathrobium geminum ; Jakovlev, 1902, Memoires de la Société des Naturalistes de Jaroslaw, 1: 111

Ryvkin A.B.

Lathrobium geminum ; Bernhauer & Schubert, 1912: 257

Lathrobium (s.str.) geminum ; Wüsthoff, 1942, Mitteilungen der Münchener Entomologischen Gesellschaft, 32: 585

Lathrobium (s.str.) geminum ; Horion, 1965: 68 Lathrobium (s.str.) geminum ; Tichomirova, 1973: 178

Lathrobium geminum ; Bordoni, 1980, Frustula Entomologica (Nuova Serie), 2(15): 20 Lathrobium geminum ; Ryvkin, 1984, Zhivotnyi mir yuzhnoy taygi: 123

Lathrobium (s.str.) geminum ; Coiffait, 1982, Nouvelle Revue d’Entomologie. Suppl., 12(4): 362 Lathrobium geminum ; Tichomirova, 1982, Pochvennyie bespozvonochnyie Moskovskoy oblasti: 215

Lathrobium geminum ; Babenko, 1982, Poleznyie i vrednyie nasekomyie Sibiri: 56

Lathrobium geminum ; Veselova & Ryvkin, 1991, Biological resources and Biocenoses of Yenissey Taiga: 192

Lathrobium geminum ; Herman, 2004, Bulletin of Zoological Nomenclature, 61(1): 25

Lathrobium geminum [?]; Shavrin, 2007a, Trudy Pribaikal’skogo natsional’nogo parka, 2: 137 Lathrobium (s.str.) geminum ; Shavrin, 2007b, Baltic Journal of Coleopterology, 7(2): 174 Lathrobium bicolor Heer, 1839: 240 Lathrobium bicolor [?]; Iljin, 1926, Revue Russe Entomologique, 19(3–4): 225

Lathrobium boreale Hochhuth, 1851 , Bulletin de la Société Impériale des Naturalistes de Moscou, 24(3): 41

Lathrobium boreale ; Hochhuth, 1863, Bulletin de la Société Impériale des Naturalistes de Moscou, 35(3): 71

Lathrobium boreale ; J. Sahlberg, 1876, Acta Societatis pro Fauna et Flora Fennica, 1: 39 Lathrobium boreale ; J. Sahlberg, 1880, Kongliga Svenska Vetenskaps-Akademiens Handlingar, 17(4): 74

Lathrobium boreale ; J. Sahlberg, 1899, Horae Societatis Entomologicae Rossicae, 32: 340 Lathrobium boreale ; J. Sahlberg, 1900, Acta Societatis pro Fauna et Flora Fennica, 19(4): 27 Lathrobium (s.str.) boreale ; Gusarov, 1992, Revue Russe Entomologique, 71(4): 784 Lathrobium boreale ; Shavrin, 2001, Proceedings of the Biology and Soil Department of the Irkutsk State University, 5: 86

Lathrobium volgense Hochhuth, 1851 , Bulletin de la Société Impériale des Naturalistes de Moscou, 24(3): 42

Lathrobium (Lathrobium) volgense ; Jakobson, 1909: 492

Lathrobium (s.str.) volgense ; Boháč, 1986, Acta entomologica bohemoslovaca, 83: 380 Lathrobium volgense ; Shavrin et al., 1999, Proceedings of the Biology and Soil Department of the Irkutsk State University, 1: 35 Lathrobium volgense ; Shavrin et al., 2001, Proceedings of the Biology and Soil Department of the Irkutsk State University, 5: 103 Lathrobium boreale Thomson, 1860: 198 Lathrobium rufescens Motschulsky, 1860 , Bulletin de la Société Impériale des Naturalistes de Moscou, 33(2): 563

Lathrobium obscuriceps Motschulsky, 1860 , Bulletin de la Société Impériale des Naturalistes de Moscou, 33(2): 564

Lathrobium castaneipenne [?]; Poppius, 1908, Acta Societatis pro Fauna et Flora Fennica, 31(6): 12

Lathrobium (s.str.) difficile Coiffait, 1953 , Annales de la Société Entomologique de France, 122: 104

Lathrobium (s.str.) fallaciosum Coiffait, 1953 , Annales de la Société Entomologique de France, 122: 104

Material. RUSSIA: CHELYABINSK AREA: 2 females ( ZMMU): Chelyabinsk Area, 20 km S Asha, nr Viliay. Old Quercus + Tilia forest with Acer & Ulmus . 15.06.1991. S.Golovatch, L.Penev, A.Vasilev, et al. leg.— K HANTY- M ANSI AUTONOMOUS REGION: 1 female ( AR): Khanty- Mansiyskiy District, Priobskoye oilfield, 760th km of road Khanty-Mansiysk–Nefteyugansk, 60°51’58,0'’N, 71°27’41,0'’E. Control. Under plants in young pinery with spots of Cladonia spp. , Cetraria islandica , Polytrichum piliferum , Pleurozium schreberi , close cover of leaf rosettes of Hieracium sp. pr. pilosella, Vaccinium vitis-idaea around rotten stubs, on SW slope of hill (xerosere). 15.08.2005. V.B.Semenov leg.—

TYUMEN AREA: 3 females ( AR) : Yarkovskiy District, 70 km S of Tobolsk, Mazurovo.

On new and poorly known Lathrobium (s.str.) species from Siberia and the Russian Far East ...

29.06.1982. N.Poryadina leg.— 1 male ( AR): same locality and collector. Birch forest. 01.07.1982.— 2 females ( AR): same locality and collector. Spruce forest. 02.07.1985.— 1 male ( AR): same locality, biotope and collector. 07.07.1985.— KRASNOYARSK TERRITORY: 1 male, 1 female ( AR): Turukhanskiy District, upper reaches of Nizhnyaya Lebedyanka River, river bank with forbs ( Urtica sp. , Trollius asiaticus , Caltha palustris , Filipendula ulmaria ): dead grass, mosses and birch-spruce litter. 18.06.1992. V.B.Semenov leg.— 1 female ( AR): same locality and collector, mosses and litter in forest with Pinus sibirica , Abies sibirica , admixture of Betula sp. , Hylocomium splendens , Pleurozium schreberi , Polytrichum commune . 19.06.1992.— 1 male, 2 females ( AR): same locality and collector, on tussocks with last year’s grass in swampy forest with Betula sp. , Abies sibirica , undergrowth of Pinus sibirica , Picea obovata and Larix sibirica . 25.06.1992.— 1 female ( AR): Turukhanskiy District, near Bor, 40 m a.s.l., in leaf litter on the bottom of dried lake channel with Rosa sp. , Padus sp. , Ribes sp. in forest with Picea obovata , Pinus sylvestris , P. sibirica , Betula pendula , Abies sibirica , Larix sibirica . 13.06.1993. V.B.Semenov leg.— 1 female ( AR): Turukhanskiy District, Kamennyi Dubches River, 60 km below Teulches River mouth, near hut. 150 m a.s.l., Picea forest with admixture of Pinus sibirica and Betula sp. : in Polytrichum commune , Pleurozium , Hylocomium . 24.09.1992. V.B.Semenov leg.— 1 male ( AR): Turukhanskiy District, Central Siberian Biosphere Reserve, Bol’shaya Varlamovka River basin, middle reaches of Bol’shaya Raskosaya River, drift of straw and needles at river bank. 23.05.1992. V.B.Semenov leg.— 2 males, 2 females ( AR): same locality and collector, edge of swamp with Sphagnum spp. , Pleurozium schreberi and Hylocomium splendens among Picea obovata and Betula sp. 25.05.1992.— 1 male ( AR): same locality and collector, Pleurozium schreberi and soil near roots of Betula sp. , Picea obovata and spruce stumps. 29.05.1992.— 1 male, 1 female ( AR): same locality and collector, moss and litter on glade with tree undergrowth ( Picea obovata , Betula sp. , Populus tremula ,

Pleurozium schreberi , Polytrichum commune , Sphagnum spp. etc.). 07.06.1992.— 2 males ( ZMMU): Krasnoyarsk Territory, Manskiy District, ‘ Stolby’ Nature Reserve , Kandalak cordon, dried mouth of Kandalak Rill , in leaf litter of willow bush with Padus sp. , Urtica sp. & Ribes nigrum . 24.08.1994. I.A.Ushakov leg.— EVENKIA AUTONOMOUS REGION: 1 male, 1 female ( AR): Baykitskiy District , Central Siberian Biosphere Reserve , Stolbovaya River basin: Birapchana River , 3–5 km below confluence of the Levaya Birapchana and Pravaya Birapchana rivers, 480 m a.s.l., river bank. 28.06.1988. A.B.Ryvkin leg.— 1 female ( AR): Baykitskiy District , Central Siberian Biosphere Reserve , Stolbovaya River : 10 km below Dulkuma River mouth, 70 m a.s.l., under great stones on river bank. 04.07.1993. V. B.Semenov leg.— IRK UTSK AREA: 1 female (AVSh): Nizhneudinskiy District , 2 km E of Yaga. 30.06.1999. A.Shavrin leg. [‘ Lathrobium brunnipes ’]— 1 female (AVSh): Tayshetskiy District , Biryusa River valley near Patrikha. 24.06.1998. A.Shavrin leg.— 1 female (AVSh): Tayshetskiy District , Biryusa River valley 5 km S Trakt-Uzhet. 03– 04.07.1999. A.Shavrin leg. [‘ Lathrobium elongatum ’]— 1 male, 2 females (AVSh): Nizhneudinskiy District , 5 km S of Vodopadnyi. 22– 23.06.1998. A.Shavrin leg. [‘ Lathrobium (s.str.) geminum Kr. det. Shavrin A. 99’]— 1 male ( AR): Zhigalovskiy District , pipeline route Kovykta-Zhigalovo , plot #2, field. 10.08.2006. V. B.Semenov & L.B.Rybalov leg.— 1 male, 1 female (AVSh): Zalarinskiy Distr. , near Bazhir. 20– 24.06.1997. A.Shavrin leg.— 4 males, 3 females (AVSh): NE of Shumilovo, N 56°02.969`E 103°17.437`, 493 m a.s.l., Abies sibirica & Pinus sibirica forest with Milium effusum L. 24– 25.06.2008. A.Shavrin & I.Enushchenko leg.— 2 males, 4 females (AVSh): 15th km of Kirensk-Podvoloshino road, left side of Lena River , N 57°49.309`E 108°17.019, 252 m a.s.l., flood-plain meadow with Bromopsis inermis & B. sibirica , and sedgy bottom of dried ephemeral reservoir at flood-plain meadow near edge of spruce-birch forest. 04– 09.08.2008. A.Shavrin & I.Enushchenko leg.— 1 male (AVSh): Khomutovo, Urik River . 25.05.2001. A. V. Shavrin leg.— 1 male (AVSh): Ust’ - Kutskiy District , Bol’shaya Tura River valley . 10.08.2008. A.Shavrin & I.Enushchenko leg.— 1 male, 1 female (AVSh):

Ust’-Kutskiy District, Nizhnyaya Bochakta River valley. 11.08.2008. A.Shavrin & I.Enushchenko

Ryvkin A.B.

leg.— 1 male (AVSh): Usol’skiy District, Toysuk River valley , 5 km S of Tal’yany. 27.07.1998. A.Shavrin leg. — 1 male (AVSh): Cheremkhovskiy District, Sredniy Bulay. 01– 02.07.2006. A. V. Shavrin leg. [‘ Lathrobium elongatum Shavrin A. det. 2006’]— 1 female (AVSh): Cheremkhovskiy District, Sredniy Bulay. 01.07.2006. A. V. Shavrin leg. — 1 female (AVSh): Alarskiy District, near Alar’ . 28.06.1997. A.Shavrin leg. [‘ Lathrobium elongatum L. Shavrin A. det. 97’]— 1 male (AVSh): Olkha River valley, Olkha. 15.09.2006. A. V. Shavrin leg. [‘ Lathrobium geminum Kr. Shavrin A. det. 2006’]— 2 females (AVSh): near Shelekhov. 06.05.1997. A.Shavrin leg. [‘ Lathrobium geminum Kr. Shavrin det. 97’]— 1 male, 1 female (AVSh): near Irkutsk, Dzerzhinsk. 18– 20.04.2007. D.Fominykh leg. — 1 female (AVSh): Angarsk, near Yelovskoye Reservoir. 21.06.2008. I. V. Enushchenko leg.

I have this species in my regular collection also from RUSSIA: PERM AREA (Gornozavodskiy District, Basegi Nature Reserve), KARELIA REPUBLIC (Kivach Nature Reserve), TATAR REPUBLIC (Volzhsko-Kamskiy Nature Reserve, Atabayevo ca. 50 km S Kazan), KOSTROMA AREA (Manturovskiy District, nr. Shilovo)[see Ryvkin 1984], MOSCOW AREA (Solnechnogorskiy District, Peshki; Podol’skiy District, Malinskoye Forestry; Podol’skiy District, Sharapovo ca. 50 km SW Moscow, nr. Krasnaya Pakhra; Istrinskiy District, nr. Pavlovskaya Sloboda; Domodedovskiy District, state farm ‘Krasnyi Put’‘; Odintsovskiy District, Zhavoronki; Moscow, Rublyovo), TULA AREA (Shchiokinskiy District, NE Krapivna, Tulskiye Zaseki Nature Reserve)— BELGOROD AREA (Belgorod), UKRAINE: TRANSCARPATIAN AREA (Mukachevo), VOROSHILOVGRAD (= LUGANSK) AREA (Stanichno-Luganskiy Nature Reserve, nr. Kondrashevka), BYELORUSSIA: VITEBSK AREA (Berezinskiy Nature Biosphere Reserve, Kraitsy), GOMEL AREA (Struki ca. 12 km NE Buda-Koshelevo).

Remarks. The species very common and widespread in Palaearctic, with long and confused taxonomic background I think to be irrelevant to discuss herein (for detailes see

Herman 2004). Its range was defined as ‘Europe and the Caucasus’ ( Bernhauer & Schubert 1912; Boháč, 1986), or ‘Europe […], manque dans les parties les plus chaudes de la région méditerranéenne’ ( Coiffait 1982), to the whole Palaearctic ( Horion 1965; Tichomirova 1973). In Siberia and the Far East, the species has been reported for the Sverdlovsk Area, Khanty-Mansi Autonomous Region, Tyumen Area, and Novosibirsk Area (J. Sahlberg 1880), Krasnoyarsk Territory (J. Sahlberg 1880; Mäklin 1881; Veselova & Ryvkin 1991), Taymyr Autonomous Region (J. Sahlberg 1880), Irkutsk Area (Shavrin 2001, 2007a; Shavrin et. al. 2001;), Maritime Province ( Horion 1965). The material listed above proves the species to be represented in some additional territories. But at the same time some earlier records seem to be open to question. I don’t know this species both from the Caucasus and from the Far East. The Far Eastern record is based on the mentioned by Horion (l.c.) four specimens labelled as ‘Wladiwostok, Frieb leg.’ (C.Koch det.). But I have seen no material on L. geminum from the territories eastwards off the Baikal Lake. Wide territories of NE Siberia from at least Central Yakutia to the Kamchatka Peninsula are also uninhabited with this species.

Figures

Figures 1–7. ― Male structures of Lathrobium spp. 1–4. L. generosum sp.n. (1–2: HT, 3–4: PT): aedeagus (1: apical part laterally, 3: laterally, 2: operculum from the side opposite to the basal opening), abdominal sternite 8 ventrally (4). 5–7. L. geminum Kraatz, 1857. (Tyumen Area:Mazurovo): aedeagus (5: apical part laterally, 6: operculum from the side opposite to the basal opening), abdominal sternite 8 ventrally (7). The long axes of spicule rows in the male abdominal sternites are shown as dash-lines. Scale = 0.1 mm.