CERATOBATRACHINAE, Boulenger, 1884

Brown, Rafe M., Siler, Cameron D., Richards, Stephen J., Diesmos, Arvin C. & Cannatella, David C., 2015, Multilocus phylogeny and a new classification for Southeast Asian and Melanesian forest frogs (family Ceratobatrachidae), Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 174 (1), pp. 130-168 : 143

CERATOBATRACHINAE BOULENGER, 1884

Diagnosis

Other species of Ceratobatrachinae differ from Alcalus by having (1) a relatively narrow head (vs. wide); and (2) smooth skin, with or without dermal tubercles and/ or dermal ridges (vs. coarsely textured, shagreened, or ‘wrinkled’ in appearance); (3) rounded terminal toe discs (vs. spatulate toe discs with squarish terminal shape); (4) absence of nuptial pads in males (vs. presence); (5) presence of a medial subgular vocal sac in most species (vs. absence); (6) presence of palmar supernumerary tubercles (vs. absence); and (7) presence of rounded subarticular tubercles and outer metatarsal tubercles on plantar surface (vs. presence, elongate). Finally, most species of Ceratobatrachinae (except Cornufer guppyi , species of the subgenus Potamorana , and Platymantis levigatus ) differ from Alcalus by the absence of interdigital webbing on the feet (vs. presence); and by having terrestrial or arboreal microhabitat preferences (vs. semiaquatic). Morphological synapomorphies for Ceratobatrachinae have not been identified.

Content

The genera Cornufer and Platymantis (see below).

Comment

We have not converted Ceratobatrachinae (Clade D) to a clade name, but have instead coined a new unranked clade name, Anurajen. If Ceratobatrachinae were converted to a clade name, and if the family Ceratobatrachidae (Node B) were re-ranked as Ceratobatrachinae in the future, then the phylogenetic name Ceratobatrachinae and the ranked name Ceratobatrachinae would refer to different clades, which would cause confusion. Therefore, we define a new clade name below denoting the same clade as the subfamily Ceratobatrachinae .