Penthoscapha toxopei (Gressitt & Sedlacek) Gressitt & Sedlacek, 2009

Riedel, Alexander, 2009, Revision of the genus Penthoscapha Heller (Coleoptera, Curculionoidea, Entiminae, Eupholini) with notes on the genera of Eupholini from New Guinea, Zootaxa 2224, pp. 1-29 : 24-25

Penthoscapha toxopei (Gressitt & Sedlacek) , comb. n.

( Figs. 16–19 , 26 , 40, 53, 60, 62, 72)

Gymnopholus toxopei Gressitt & Sedlacek, 1967: 486 (type locality: “Mist camp”).

Diagnosis. Body subglabrous, with sparse scattered setae, but without squamose patches. Rostrum relatively short (Fig. 40). Elytral intervals in basal half markedly transversely rugose, even, without costae; sutural interval subapically with rounded protrusion ( Fig. 19 ).

Description. Paratype, female ( Figs. 16–17 ). Length. Total length 24.0 mm; pronotum + elytron 20.0 mm.

Coloration black; without exocuticular pigments, showing traces of aging, partly covered with dull deposits, (these completely missing in holotype with cleaner, more shining surface) partly shining; subglabrous, nude except for recumbent inconspicuous setae and scattered bluish scales on legs and antenna.

Head dull, subglabrous; between eyes 1.07 X as wide as vertical diameter of eye, bordering eyes with shallow punctures, medially with small fovea. Vertex simple, without median furrow. Eyes moderately prominent from outline of head. Gena subequal (0.98 X) to vertical diameter of eye.

Rostrum (Fig. 40) distinctly shorter than pronotum [0.75 X; in conflict with information in key of Gressitt & Sedlacek, 1967: 485]; 1.70 X as long as wide at base; maximum width in front of antennal insertion 1.40 X width at base. Dorsum at middle weakly concave, but without distinct median furrow; in front of eyes with distinct sublateral impressions.

Antenna. Funicle + club 1.70 X length of scape; club 3.06 X as long as wide.

Thorax. Pronotum ca. as long as wide [1.02 X wider than long], sides gently sinuate, widest in apical half; disc transversely rugose, with shallow impression slightly in front of middle, midline weakly impressed. Thoracic venter subglabrous.

Elytron medially free, 3.94 X longer than wide between humeri; at greatest width 1.15 X wider than between humeri; side evenly rounded; dull, coriaceous; striae regular, distinct, moderately impressed; intervals markedly transversely rugose, especially in basal half, intervals 3 and 7 simple, not forming longitudinal ridges; sutural interval of elytron at apical declivity ( Fig. 19 ) forming rounded protrusion visible in lateral aspect. Ala ( Fig. 26 ) short, length 17.5 mm, 1.31 X length of elytron.

Legs. Femora moderately long; metafemur in repose not reaching elytral apex. Tibiae ventrally subapically sparsely denticulate; metatibia apically with sparsely squamose bevel; posterior distal comb of metatibia consisting of few dentiform setae, terminating ventrally on a protuberance (“secondary mucro”). Tarsi elongate, tarsomere 1 of metatarsus 2.4 X as long as wide, tarsomere 2 ca. as long as wide.

Abdomen. Venter subglabrous, weakly punctate, with sparse recumbent setae. Terminalia. Fig. 60 . Tergite VIII moderately narrow, sides converging to pointed apex. Hemisternite ( Fig. 53 ) 2.2 X as long as high. Stylus 3.3 X as long as wide, subapically with dense cluster of short stiff setae pointing laterad. Spermatheca ( Fig. 62 ) with gland ca. 3 X as long as body.

Material examined. Holotype, female; only three photographs were available ( BPBM). Paratype, female: WEST NEW GUINEA, Jayawijaya Reg.: labels ( Fig. 18 ): 1) [white, rectangular]: “Neth. Ind.- American New Guinea Exped., Mist Camp [E 139° 4.2´S 03° 33.8´], 1800 m, I.1939, L.J.Toxopeus [printed]”, 2) [yellow, rectangular] “ Paratype [printed] Gymnopholus toxopei [handwritten] Gressitt & [printed] Sedl. [handwritten]; 3) [red, rectangular] “ Paratype [printed]” ( RMNH).

Distribution ( Fig. 72 ). Jayawijaya Reg. (Mist camp). Elevation: 1700-1800 m. So far, known from the two type specimens only. The “mist camp” is synonymous with the “1,800-meter camp” of the 1938-1939 Archbold expedition and was located “ 15 km southwest of Bernhard camp” ( Archbold et al., 1942); the latter was located on the Idenburg River. Based on these data, the map of Archbold et al. (1942), and Google Earth the approximate coordinates of the type locality are E 139° 4.2´S 03° 33.8´. The elevation is relatively low for this group of usually subalpine weevils. However, since the mountain where it was collected rises steeply from the lowlands, the zonation of the vegetation is somewhat depressed compared to other highland regions.

Figures

FIGURES 16 – 19. Penthoscapha toxopei, paratype (16) dorsal aspect; (17) lateral aspect (18) labels; (19) elytral apex, lateral aspect.

FIGURES 26 – 31. Left wing of Penthoscapha species: (26) P. t o x o p e i, paratype; (27) P. similis, holotype; (28) P. doormanensis, holotype; (29) P. p u l v ere a, holotype; (30) P. gerhardschereri, holotype; (31) P. l o re n t z i, lectotype.

FIGURES 60 – 66. Female terminalia of Penthoscapha species: (60) P. t o x o p e i, paratype; (61) P. similis, allotype; (62) P. toxopei, paratype, spermatheca; (63) P. gerhardschereri, allotype, spermatheca; (64) P. similis, allotype, spermatheca; (65) P. gerhardschereri, allotype; tergite VIII and ductus spermathecae; (66) P. similis, allotype, tergite VIII.

FIGURES 53 – 59. Female terminalia of Penthoscapha species: (53) hemisternite of P. toxopei, paratype; (54) hemisternite of P. gerhardschereri, allotype; (55) hemisternite of P. similis, allotype; (56) tergite VII of P. gerhardschereri, allotype; arrow marking wing-folding patch; (57) tergite VII of P. similis, allotype; arrow marking wing-folding patch; (58) sternite VIII of P. gerhardschereri, paratype; (59) sternite VIII of P. similis, allotype.

FIGURES 71 – 72. Distribution of Penthoscapha species.