Darkoneta, Ledford, Joel M. & Griswold, Charles E., 2010

Ledford, Joel M. & Griswold, Charles E., 2010, A study of the subfamily Archoleptonetinae (Araneae, Leptonetidae) with a review of the morphology and relationships for the Leptonetidae, Zootaxa 2391, pp. 1-32 : 14-16

publication ID	https://doi.org/ 10.5281/zenodo.193866
DOI	https://doi.org/10.5281/zenodo.6204730
persistent identifier	https://treatment.plazi.org/id/881A879D-FF97-FFDC-FF5D-F9631E61FCB7
treatment provided by	Plazi
scientific name	Darkoneta
status	gen. nov.

Treatment

Darkoneta , gen. nov.

( Figs. 11–17 View FIGURES 11 – 17 , 29, 30, 32, 34 View FIGURES 29 – 34 , 46 View FIGURES 43 – 48 , 57–64 View FIGURES 57 – 64 , 79–84 View FIGURES 79 – 84 , 85–90 View FIGURES 85 – 90 , 91–96 View FIGURES 91 – 96 , 111, 112 View FIGURES 109 – 114 , 116 View FIGURES 115 – 116 )

Type species. Archoleptoneta stridulans Platnick, 1994 , here designated.

Etymology. This genus is named in honor of Darko Ljubić (Darrell Ubick), a trusted friend, colleague, and collector of many leptonetids throughout North America and Europe.

Diagnosis. Small (0.87-1.28 mm), three-clawed, ecribellate, six-eyed or blind spiders separated from all other leptonetids except Archoleptoneta by having the eyes contiguous ( Figs. 11–13 View FIGURES 11 – 17 ); male palpal tarsus cylindrical; bulb suboval with three accessory sclerites and a tapering embolus ( Figs. 80, 83 View FIGURES 79 – 84 , 86, 89 View FIGURES 85 – 90 , 92, 95 View FIGURES 91 – 96 ); females with simple, paired receptacula ( Fig. 111–112 View FIGURES 109 – 114 ). Separated from Archoleptoneta by having a rectangular colulus ( Fig. 46 View FIGURES 43 – 48 , 57, 61 View FIGURES 57 – 64 ) and males with a stout prolateral accessory sclerite (PRS, Figs. 80, 83 View FIGURES 79 – 84 , 86, 89 View FIGURES 85 – 90 , 92, 94 View FIGURES 91 – 96 ).

Synapomorphies. Darkoneta gen. nov. species are united by the presence of a colulus ( Figs. 46 View FIGURES 43 – 48 , 57, 61 View FIGURES 57 – 64 ) and an enlarged, modified prolateral sclerite on the male palpal bulb (PRS, Figs. 80, 83 View FIGURES 79 – 84 , 86, 89 View FIGURES 85 – 90 , 92, 94 View FIGURES 91 – 96 ).

Description. Total length 0.87–1.28. Carapace pale to orange, round to oval in dorsal view ( Figs. 6–8 View FIGURES 6 – 10 , 16 View FIGURES 11 – 17 ), elevated in lateral profile ( Fig. 16–17 View FIGURES 11 – 17 ), length 1.10–1.33 x width; smooth; thoracic fovea inconspicuous; carapace rounded along posterior margin in females, males of D. stridulans and D. quetzal sp. nov. with posterior stridulatory structures ( Fig. 12–13 View FIGURES 11 – 17 , 29–30 View FIGURES 29 – 34 ), all other species unmodified; six eyes ( Fig. 29 View FIGURES 29 – 34 ) or blind ( Fig. 32 View FIGURES 29 – 34 ), AME absent; ALE and PLE with dark to dusky markings ( Fig. 11–13 View FIGURES 11 – 17 ); OA wider than long, with a single seta anteriad of ALE, two setae posteriad of PLE; fang furrow with 5–6 teeth on the promargin and 0–2 teeth on the retromargin. Sternum smooth, tapering posteriorly, as long as wide. Abdomen ( Figs. 11–12, 14– 17 View FIGURES 11 – 17 ) oval, pale to light-brown, covered with fine setae; stridulatory files present anteriorly in males of D. stridulans and D. quetzal sp. nov.; spinning organs ( Figs. 46 View FIGURES 43 – 48 , 57–64 View FIGURES 57 – 64 ); colulus recatangular ( Fig. 46 View FIGURES 43 – 48 , 57, 61 View FIGURES 57 – 64 ); ALS cylindrical, with 1–3 tartipores, a single elongate spigot anteriorly, and 4 spigots of varying size on short, circular bases, and at least one spigot lacking a base; PMS tetrahedral, females with 2–4 spigots on short, circular bases, 1–2 spigots on a thickened base anteriorly ( Fig. 59 View FIGURES 57 – 64 ), with or without a single nubbin ( Fig. 59 View FIGURES 57 – 64 ), male PMS with 1–2 spigots on short, circular bases; PLS with 1–2 spigots on an elongate pointed base in females, absent in males. Leg formula I, IV, II, III or IV, I, II, III; legs elongate and thin, sparsely covered with fine setae and lacking spines; male femur I 1.19–2.80 x carapace width, females 1.13–2.3 x carapace width; middorsal integumentary glands elongate, with single, small pores ( Figs. 7–8 View FIGURES 6 – 10 in Platnick, 1994). Male palpal femur length 0.50–0.78 x carapace width. Basal haematodocha expandable, bulb suboval, bearing three flexibly attached accessory sclerites at base ( Figs. 80, 83 View FIGURES 79 – 84 , 86, 89 View FIGURES 85 – 90 , 92, 95 View FIGURES 91 – 96 ), embolus tapered or broad. Female genitalia ( Fig. 111–112 View FIGURES 109 – 114 ) haplogyne, consisting of simple, paired recaptacula.