Archoleptoneta Gertsch, 1974

Ledford, Joel M. & Griswold, Charles E., 2010, A study of the subfamily Archoleptonetinae (Araneae, Leptonetidae) with a review of the morphology and relationships for the Leptonetidae, Zootaxa 2391, pp. 1-32 : 10-11

publication ID	https://doi.org/ 10.5281/zenodo.193866
DOI	https://doi.org/10.5281/zenodo.6204720
persistent identifier	https://treatment.plazi.org/id/881A879D-FF93-FFD9-FF5D-F84B183FFA94
treatment provided by	Plazi
scientific name	Archoleptoneta Gertsch, 1974
status

Treatment

Genus Archoleptoneta Gertsch, 1974

Type species. Archoleptoneta schusteri, Gertsch 1974 .

Archoleptoneta Gertsch 1974: 198 (type species Archoleptoneta schusteri Gertsch 1974: 199 ); Platnick 2008. Archoleptoneta (part): Brignoli 1974: 210 –212; Brignoli 1977: 215 –217; Ledford 2004: 122; Platnick 1994.

Diagnosis. Small (1.16–1.82 mm), three-clawed, cribellate, six-eyed spiders separated from other leptonetids except Darkoneta gen. nov. by having the eyes contiguous ( Figs. 6–8 View FIGURES 6 – 10 , 23, 25 View FIGURES 23 – 28 ); male palpal tarsus cylindrical, lacking a dorsal groove ( Fig. 78 View FIGURES 73 – 78 ); bulb suboval with three accessory sclerites ( Figs. 76–77 View FIGURES 73 – 78 , 98, 101 View FIGURES 97 – 102 , 104, 107 View FIGURES 103 – 108 ); females with simple, paired receptacula ( Figs. 109 View FIGURES 109 – 114 ). Separated from Darkoneta gen. nov. by having a cribellum, males with a long, tapering embolus that extends to the tip of the tarsus ( Fig. 76–78 View FIGURES 73 – 78 , 98, 99 View FIGURES 97 – 102 , 104, 105, 108 View FIGURES 103 – 108 ), and a straight, unmodified prolateral sclerite (PRS, Figs. 98–99, 101–102 View FIGURES 97 – 102 , 104–105, 107–108 View FIGURES 103 – 108 ). Females with spermathecae that are longer than wide ( Figs. 109–110 View FIGURES 109 – 114 ).

Synapomorphies. Archoleptoneta species are united by features of the male palpal bulb, including the presence of a straight prolateral sclerite (PRS, Figs. 98, 101–102 View FIGURES 97 – 102 , 104–105, 107 View FIGURES 103 – 108 ) and a straight median sclerite (MS, Figs. 98, 101 View FIGURES 97 – 102 , 104, 107 View FIGURES 103 – 108 ).

Description. Total length 1.16–1.82. Carapace pale to light-brown, round to oval in dorsal view ( Figs. 6–8 View FIGURES 6 – 10 , 23 View FIGURES 23 – 28 ), length 1.14–1.30 x width; smooth; thoracic fovea inconspicuous; rounded along posterior margin in both sexes; six eyes, AME absent; ALE and PME with dark to dusky markings, u-shaped at posterior margin ( Figs. 7 View FIGURES 6 – 10 , 25 View FIGURES 23 – 28 ); OA wider than long, with a single seta anteriad of ALE, two setae posteriad of PLE; chelicerae unmodified, fang promargin with a single large proximal tooth and 6–8 denticles. Sternum smooth, oval, tapering posteriorly, as long as wide. Abdomen ( Figs. 6, 8 View FIGURES 6 – 10 ) oval, pale to light-brown, covered with fine setae; spinning organs ( Figs. 43–44, 47 View FIGURES 43 – 48 , 49–56 View FIGURES 49 – 56 ); cribellum oval, divided, with uniformly distributed strobilate spigots, reduced in males ( Fig. 53 View FIGURES 49 – 56 ); ALS cylindrical, with 3–4 tartipores, with at least two major ampullate gland spigots and 3–5 piriform gland spigots; PMS tetrahedral, females with 2 anterior cylindrical gland spigots, 2 aciniform gland spigots posteriorly ( Fig. 51 View FIGURES 49 – 56 ), males retain 2 aciniform gland spigots; female PLS with 2 posterior cylindrical gland spigots and a single spigot on an elongate pointed base, which is replaced by a nubbin in males. Leg formula IV, I, II, III or I, IV, II, III; legs elongate and thin, sparsely covered with fine setae and lacking spines; male femur I 1.50–1.90 x carapace width, females 1.50–1.89 x carapace width; middorsal integumentary glands elongate, with single, small pores ( Fig. 31 View FIGURES 29 – 34 ). Male palpal femur length 0.57– 0.77 x carapace width; palpal tarsus cylindrical in dorsal view ( Fig. 78 View FIGURES 73 – 78 ). Basal haematodocha expandable, bulb suboval, bearing three ventral accessory sclerites at base ( Figs. 76 View FIGURES 73 – 78 , 98, 101 View FIGURES 97 – 102 , 104, 107 View FIGURES 103 – 108 ) embolus straight to sinuous, extending to the tip of the tarsus ( Fig. 76 View FIGURES 73 – 78 , 98 View FIGURES 97 – 102 , 104–105, 108 View FIGURES 103 – 108 ); tarsal organ trilobed ( Figs. 37, 39 View FIGURES 35 – 42 ). Female genitalia ( Figs. 109–110 View FIGURES 109 – 114 ) haplogyne, consisting of simple, paired spermathecae.