Tragardhus (Mitragardhus) ryszardi, Kamiński, 2017

Kamiński, Marcin Jan, 2017, Afrotropical Melambiina: revision of the genus Tragardhus Koch, 1956 and redescription of Pseudemmallus Koch, 1956 (Tenebrionidae: Pedinini), Zootaxa 4347 (1), pp. 71-93 : 75-76

Tragardhus (Mitragardhus) ryszardi sp. nov.

http://zoobANk.oRg/URN:lsiD:zoobANk.oRg:Act:29C8C4A5-6730-497D-B73E-C172DA362BC4 ( Figs 3C, E , 7 )

Type data. Holotype, mAle (TMNH): “Z.A. 79 / AlfReD Dist. / ORibi GoRge”, “HUmUs / X-1961 ”, “N. LeleUp leg” ( Fig. 3C ).

Etymology. NAmeD AfteR RYszARD RAś (boRN oN JUNe 12th, 2015), soN of mY fRieNDs MAgDAleNA WoźNiAk AND MARciN RAś.

Diagnosis. This species is similAR to T. (N.) nodosus bY hAviNg eloNgAteD tUbeRcles oN 3RD, 5th AND 7th elYtRAl iNteRvAls. Tragardhus (M.) ryszardi cAN be eAsilY DistiNgUisheD bY hAviNg sleNDeR AND glAbRoUs metAtibiAe iN mAles (wiDeNeD AND coveReD bY A loNgitUDiNAl pAtch of setAe iN nodosus ).

Description. BoDY—leNgth cA. 3.5 mm. HAbitUs As iN Fig. 3C .

DoRsAl siDe of heAD coveReD with coNspicUoUs AND coNflUeNt pUNctURes (0.1 DiAmeteRs ApARt). FRoNtoclYpeAl sUtURe coARse; with A pAiR of Deep DepRessioNs locAteD NeAR meDiAN pARt. CANthUs sUbpARAllel At bAse. ClYpeAl emARgiNAtioN Deep (clYpeAl emARgiNAtioN wiDth: Depth RAtio cA. 4.2). LAbRUm wiDe (wiDth: leNgth RAtio cA. 3.0); DelicAtelY emARgiNAte iN miDDle; posteRioR thiRD coveReD with DeNse AND coNspicUoUs setAe. MeNtUm wiDest iN miDDle; flAt, Not coveRiNg cARDo oR stipes; ApicAllY with shAllow meDiAN iNDeNtAtioN; lAteRAl lobes, iN veNtRAl view, coveReD bY miDDle pARt. SUbmeNtUm tRiANgUlAR. ApicAl segmeNt of mAxillARY pAlpUs bRokeN off of the stUDieD specimeN. PAlpifeR with pAtch of loNg setAe locAteD NeAR bAse. EYe Not shielDeD bY ANY keel. VeNtRAl pARt of eYe ReDUceD iN RelAtioN to DoRsAl pARt; two fAcets pReseNt betweeN cANthUs AND temple. ANteNNA RelAtivelY loNg (ANteNNA: pRoNotUm leNgth RAtio cA. 1.1); thiRD ANteNNomeRe shoRt (3RD: 2ND ANteNNomeRe leNgth RAtio cA. 1.3); fifth ANteNNomeRe shoRt (leNgth: wiDth RAtio cA. 1.0); ANteNNomeRes 7–10 tRANsveRse.

PRoNotUm NARRoweR thAN elYtRA (elYtRA: pRoNotUm wiDth RAtio cA. 1.4); RelAtivelY wiDe (leNgth: wiDth RAtio of pRoNotAl Disc cA. 0.6); Disc DUll, coveReD with coNflUeNt pUNctURes, eAch poiNt eqUippeD with A siNgle setA. LAteRAl siDes of pRoNotAl Disc bisiNUAte. Disc wiDest iN miDDle; withoUt ApophYseAl DepRessioNs. LAteRAl emARgiNAtioN of pRoNotUm composeD of ADjAceNt tUbeRcles; bAsAl AND ANteRioR emARgiNAtioN AbseNt. BAse of pRoNotUm slightlY bisiNUAte. PosteRioR ANgles Not pRotRUDiNg beYoND bAse. PRoNotAl hYpomeRoN shiNY, with coARse pUNctURes (1.0 DiAmeteRs ApARt). INteRcoxAl pRocess of pRosteRNUm DepResseD ApicAllY. PRocoxAe NARRowlY sepARAteD (pRocoxA: iNteRcoxAl pRocess wiDth RAtio cA. 5.0).

ElYtRAl stRiAe Not impResseD, visible As teN Rows of coNspicUoUs pUNctURes oN eAch elYtRoN (1.0 DiAmeteR ApARt)—of which two ARe sitUAteD veNtRAllY. INteRvAls 3, 5 AND 7 coveReD with eloNgAteD tUbeRcles (most eviDeNtlY visUAliseD iN bAsAl pARt); RemAiNiNg iNteRvAls coveReD with smAll RoUNDeD tUbeRcles (1.5–2.0 DiAmeteRs ApARt) eAch eqUippeD with NoticeAble setA. ElYtRAl bAse v-like; emARgiNAte. EpipleURoN NARRowiNg towARDs Apex; slightlY NARRoweR thAN NeighboRiNg elYtRAl iNteRvAl. IN its bAsAl hAlf, 9th iNteRvAl is oN sAme plANe As epipleURoN; posteRioRlY twistiNg to A DiffeReNt plANe thAN epipleURoN. ElYtRAl hUmeRUs DeNtAte, pRotRUDiNg lAteRAD. ScUtellUm RelAtivelY smAll; tRiANgUlAR. MetAthoRAcic wiNgs AbseNt.

MesoveNtRite withoUt pRocesses. MetAveNtRite RelAtivelY shoRt (metAcoxAl cAvitY: metAveNtRite (betweeN iNseRtioNs of meso- AND metAcoxAe) leNgth RAtio cA. 2.5). PRocess of fiRst AbDomiNAl veNtRite NARRow (DistANces betweeN mesocoxAe: metAcoxAe RAtio cA. 0.7). Fifth AbDomiNAl veNtRite UNboRDeReD. BAsAl two-thiRDs coveReD bY Deep pUNctURes (1.0 DiAmeteR ApARt), sURfAce betweeN pUNctURes coveReD with micRoscUlptURe; ApicAl thiRD Not pUNctAte.

Meso- AND metAtRochANteRs NeARlY eNtiRelY coveReD bY Respective femoRA; eqUippeD with A siNgle setAe. MetAcoxA wiDeNeD (wiDth: leNgth RAtio cA. 1.4). TARsi bRokeN off of the stUDieD specimeN. MAle pRotibiA with meDiAN DilAtAtioN, coveReD with hAiR; metAtibiA sleNDeR, glAbRoUs, with shARp Apex (As iN T. (M.) nodosus — Fig. 2G ); otheR leg pARts simple. MoRphologY of femAle legs UNkNowN.

AeDeAgAl tegmeN wiDest iN its bAsAl thiRD; UNipARtite. PARAmeRes DiviDeD (Up to 0.2 of whole tegmeN leNgth). ClAvA slightlY cURveD ApicAllY (hook-like); twisteD meDiAllY (NeAR its coNNectioN with tegmeN). PeNis of eveN wiDth oN most of its leNgth. BAsAl ApophYses of peNis RelAtivelY shoRt (cA. 0.25 of whole tegmeN leNgth).

MoRphologY of femAle teRmiNAliA UNkNowN.

Distribution. This species hAs beeN collecteD iN the followiNg ecoRegioNs of SoUth AfRicA (CApe ZoNe): KwAZUlU-CApe coAstAl foRest mosAic ( Fig. 7 ).

Figures

FIGURE 2. Diagnostic characters proposed for Mitragardhus, Tragardhus and chosen species. Pronotal disc (A, B, C), epipleuron (C), 5 th abdominal ventrite (D, E), elytral disc (F, H, I), metatibia (G) and protibia (J). Tragardhus (Tragardhus) biapicalis Koch, 1956 (I), T. (T.) glandipleurum Koch, 1956 (A, E, H, J), T. (Mitragardhus) ewae sp. nov. (B, F) and T. (M.) nodosus Koch, 1956 (D, G). White arrow indicates a characteristic apex of the metatibia of T. (M.) nodosus.

FIGURE 3. Habitus images of the studied species. Tragardhus (Mitragardhus) ewae sp. nov., paratype (A), T. (M.) nodosus Koch, 1956, syntype (B), T. (M.) ryszardi sp. nov., holotype (C, E), T. (M.) zuzannae sp. nov., holotype (D, F) and Pseudemmallus aspericollis (G).

FIGURE 7. Distribution of the genus Tragardhus.