Tragardhus Koch, 1956

Kamiński, Marcin Jan, 2017, Afrotropical Melambiina: revision of the genus Tragardhus Koch, 1956 and redescription of Pseudemmallus Koch, 1956 (Tenebrionidae: Pedinini), Zootaxa 4347 (1), pp. 71-93 : 73

Genus Tragardhus Koch, 1956

Tragardhus Koch, 1956: 369 .— Kulzer 1963: 431, Iwan 2004: 739, Banaszkiewicz 2006: 65, Kamiński 2011: 321.

Type species. Tragardhus (Tragardhus) glandipleurum Koch, 1956 ; bY oRigiNAl DesigNAtioN.

Diagnosis. The followiNg chARActeR combiNAtioN is UNiqUe foR Tragardhus withiN MelAmbiiNA: eYe Not DiviDeD bY exteNDiNg geNAl cANthUs iNto DoRsAl AND veNtRAl pARts (Not As iN Oreomelasma EspAñol, 1975 ; Fig. 1F ), ANteNNA (Not As iN Hanstroemium ; Fig. 1G ) AND pRotibiA RelAtivelY sleNDeR (Not As iN e.g., Bermejoina EspAñol, 1944; Fig. 1H ), some of the elYtRAl iNteRvAls coveReD bY moRe oR less eloNgAteD tUbeRcles ( Fig. 3E, F ), lAteRAl siDe of epipleURoN withoUt elevAteD cARiNA ( Fig. 1N vs 1M), fifth AbDomiNAl veNtRite withoUt sUbmARgiNAl sUlcUs (Not As iN PAlAeARctic MelAmbiiNA; Figs 1I, J ) AND pARAmeRes of AeDeAgUs DeeplY sepARAteD iN the ApicAl pARt (Not meRgeD As iN PAlAeARctic MelAmbiiNA; e.g., Fig. 1B, C ).

Figures

FIGURE 1. Diagnostic characters of different biogeographic components of Melambiina. Aedeagal tegmen (A – C), spermatheca (D, E), head (F, G, L), protibia (H), 5 th abdominal ventrite (I, J) and elytra (K, M, N). Bermejoina aiunica Español, 1944 (E, H), Gridelliopus subsquamosus Koch, 1956 (K, L), Hanstroemium adelostomoide Koch, 1953 (G), Melambius barbarus (Erichson, 1841) (D), Melambius mideltensis Antoine, 1932 (B, J), Melasmana appenhageni Koch, 1948 (C), Oreomelasma oromii Español, 1975 (F), Peyerimhoffius plicatus (Lucas, 1846) (I), Silvestriellum alatum Koch, 1956 (A), Tragardhus ewae sp. nov. (N) and Zadenos delalandei (Mulsant & Rey, 1854) (M). Abbreviations: bc. — bursa copulatrix, spspermatheca, — sag—spermathecal accesory gland, ep—epipleuron, IX. and X in. — elytral intervals 9 and 10.

FIGURE 3. Habitus images of the studied species. Tragardhus (Mitragardhus) ewae sp. nov., paratype (A), T. (M.) nodosus Koch, 1956, syntype (B), T. (M.) ryszardi sp. nov., holotype (C, E), T. (M.) zuzannae sp. nov., holotype (D, F) and Pseudemmallus aspericollis (G).