Stegosastyrus hemiclara Pyrcz, Boyer & Zacca

Zacca, Thamara, Mielke, Olaf H. H., Pyrcz, Tomasz W., Casagrande, Mirna M., Freitas, André V. L. & Boyer, Pierre, 2013, Stegosatyrus, a new genus of Euptychiina from the grasslands of neotropical realm (Lepidoptera: Nymphalidae: Satyrinae), Zootaxa 3682 (2), pp. 331-350 : 343-346

Stegosastyrus hemiclara Pyrcz, Boyer & Zacca , n. sp.

( Figs 13–16 , 61–64 , 84 )

Type locality. Peru, department of Puno, Carcel Punco.

Diagnosis. Easily differentiated from S. imbrialis n. comb. by the smaller, single ocellus on the dorsal face of the forewing.

Description. MALE ( Figs 13–14 , 61–63 ). Head: Antenna dark brown dorsally with white scales at the base of each segment, chocolate brown ventrally with white scales, club twice as wide as shaft, same color but without scales; frons with a tuft of light grey brown hair; labial palpi covered with dark brown hair; eyes chocolate brown, smooth. Thorax: dorsally dark brown, ventrally pale brown, hairy; legs pale yellowish brown. Abdomen: dorsally dark brown, laterally and ventrally pale brown. Wings: FW (length: 18.5mm): triangular with a sub-pointed apex; dorsally varying between fallow brown and chestnut, with two dark brown postdiscal and submarginal irregular lines (not apparent in some specimens), nearly touching each other in the tornal area, and gradually fainting towards the wings' subapical area; a single rounded black subapical ocellus between M1-M2, ringed with yellow and pupilated with lilac; in some specimens, with a minute lilac spot between M2-M3; a regular dark brown marginal line running parallel to a dark brown outer margin; fringes chestnut; ventrally, the FW is a shade lighter fallow brown or chestnut than on the upperside, subapical and apical area suffused with ashy scales; in some specimens, a faint irregular brown postbasal line, along with a dark brown straight postdiscal line running from subcosta to tornus where they merge with a dark brown submarginal line also extending to the subcosta; two fine, parallel dark brown marginal lines; a large rounded subapical black ocellus between M1-M2 with two minute lilac pupils, ringed with yellow, and with a minute lilac spot on its lower section. HW: dorsally chestnut, hairy on basal third, a small black submarginal ocellus between CuA1-CuA2 with a minute double lilac pupil, ringed with yellow; a fine, dark brown submarginal line parallel to outer margin, and a slightly more prominent dark brown marginal line; fringes medium brown; ventrally, the HW is fallow brown heavily overcast with ashy grey, in some specimens liberally speckled with brown; a medium brown, faint and regular postbasal line from costa to base of CuA2 vein, in some specimens not apparent; a nearly straight, except for a small incision along M3, postdiscal dark brown line from Rs to anal margin near tornus, faint in some specimens, the area basal to this line more markedly brown due to the lighter suffusion of grey than on the rest of the wing; in three out of four specimens, a small black submarginal ocellus between M1-M2 ringed with yellow, and another, similar ocellus between CuA1-CuA2 with a double, minute lilac pupil; a series of fine medium brown submarginal and marginal lines, shaped as on the upperside. Genitalia: tegumen robust with a straight dorsal surface; uncus robust, 1½ times the length of tegumen with a blunt apex; gnathos thin, elongated, approximately the length of uncus, curved upwards, with a pointed apex; appendix angular small with a curved tip; saccus short, approximately ½ the length of gnathos; valva elongated, gradually narrowing apically, with a smooth dorsal surface, and a lightly crenulated apex; aedeagus slightly longer than valva and narrow, with anterior portion lanceolate and the dorsal posterior portion split.

FEMALE ( Figs 15–16 , 64 ): Slightly larger than the male, with more rounded wings, forewing apex blunt; a shade lighter chestnut on the upperside and considerably lighter and paler on the underside, especially on the hindwing whose chestnut ground color is almost entirely overcast with light grey scaling, postbasal line barely noticeable, postdiscal line faint, and submarginal ocelli smaller; brown submarginal line not apparent. Genitalia ( Fig. 64 ): Papillae anales 2/3 the length of tergite VIII; sternite VIII separated from tergite VIII; lamella antevaginalis sclerotized; ductus bursae weakly sclerotized; bursa copulatrix oval, signa 2/5 the length of bursa.

Distribution ( Fig. 84 ). So far, known exclusively from the type locality, Carcel Punco in southern Peru. It possibly occurs farther south to Yungas de La Paz. It was collected at a very high elevation, approximately 3,000 m.

Host plant. Unknown

Examined material. Male Holotype with the following labels: Holotypus / PERU, Puno: Limbani vers Carcel Punco km 4, 3000m, 16.X.2004, P. Boyer leg, PB/ ( MZUJ)/ Stegosatyrus hemiclara Pyrcz, Boyer & Zacca det. 2013. Alotype with the following labels: Allotypus / PERU, Puno: Limbani vers Carcel Punco km 4, 3000m, 16.X.2004, P. Boyer leg, PB/ ( MZUJ)/ Stegosatyrus hemiclara Pyrcz, Boyer & Zacca det. 2013. Paratypes with the following labels: 33 Paratypus / PERU, Puno: Limbani vers Carcel Punco km 4, 3500m, 16.X.2004, P. Boyer leg, PB/ ( MZUJ)/ Stegosatyrus hemiclara Pyrcz, Boyer & Zacca det. 2013; 13 Paratypus / PERU, Puno: Limbani vers Carcel Punco km 4, 3000m, 16.X.2004, J. Bottger leg/ ( MZUJ)/ Stegosatyrus hemiclara Pyrcz, Boyer & Zacca det. 2013.

Figures

FIGURES 84. Geographical distribution of Stegosatyrus, n. gen.

FIGURES 1 – 16. Adults of Stegosatyrus, n. gen.. 1 – 2. S. imbrialis, male: 1. Dorsal, 2. Ventral; 3 – 4. S. imbrialis, female: 3. Dorsal, 4. Ventral; 5 – 6. S. ocelloides, male: 5. Dorsal, 6. Ventral; 7 – 8. S. ocelloides, female: 7. Dorsal, 8. Ventral; 9 – 10. S. periphas, male: 9. Dorsal, 10. Ventral; 11 – 12. S. periphas, female: 11. Dorsal, 12. Ventral. 13 – 14. S. hemiclara n. sp., holotype, male: 13. Dorsal, 14. Ventral; 15 – 16. S. hemiclara n. sp., allotype, female: 15. Dorsal, 16. Ventral.

FIGURES 61 – 64. Stegosatyrus hemiclara Pyrcz, Boyer & Zacca n. sp.: 61 – 63. Male genitalia (scale = 0, 05 mm): 61. Lateral, 62. Dorsal, 63. Aedeagus; 46. Female genitalia, lateral (scale = 1 mm).