Polymorphanisus muluensis Barnard, 1980

Uy, Christine Jewel C., Malicky, Hans & Bae, Yeon Jae, 2018, Review of the filter-feeding caddisfly subfamily Macronematinae (Trichoptera: Hydropsychidae) in tropical Southeast Asia, Raffles Bulletin of Zoology 66, pp. 664-703 : 690

Polymorphanisus muluensis Barnard, 1980

( Figs. 186 , 195 , 208–210, 228)

Polymorphanisus muluensis Barnard, 1980: 87 [Holotype male; Borneo; NHMUK].

Diagnosis. Male. Antennae 50 mm in length. Antennal flagellum brown with darker striations. Forewing 15 mm and hind wing 10 mm in length. Mesoscutum without evident markings. Mesoscutellum with paired dark brown markings as shown ( Fig. 186 ). Tibial spurs 2.3.3.

Male genitalia. Inferior appendages two-segmented, harpago blunt and shorter than in other species. In dorsal view, phallus with distinctive wide opening because of phallotremal sclerite (Fig. 210). Segment X divided medially, each half with double lobular structure ( Fig. 228 ).

Female. Antennae 45 mm in length. Antennal flagellum brown with darker striations. Forewing 20 mm and hind wing 15 mm in length. Mesoscutum and mesoscutellum same as male ( Fig. 186 ). Tibial spurs 2.3.3.

Material examined. Vietnam: 2 males, Nam Cat Tien, coll. H. Malicky, 17–25 June 1995 (HMPC) ; Laos: 2 females, Central Province, Viangchan, coll. Holzschuh, 1–8 June 1996 (HMPC) ; Thailand: 1 male, Sakon Nakhon Province, Nam Phung, coll. C. J. Uy, 20 May 2015 ( UPLBMNH) .

Distribution. Borneo, Laos, Thailand, Vietnam.

Remarks. Barnard (1980) provided a detailed redescription of this species. However, we examined individuals with different tibial spur formula of 1.3.2 or 1.3.3. Other diagnostic characters are illustrated in Figs. 186 , 195 , 208–210, and 228.

Figures

Figs. 183–192. Polymorphanisus thorax. 183, astictus, ocularis male and female thorax; 16, fuscus male thorax; 184–185, fuscus female thorax; 186, muluensis male and female thorax; 187, nigricornis male and female thorax; 188, quadripunctatus male and female thorax; 189, semperi male and female thorax; 190, scutellatus male thorax; 191, scutellatus female thorax; 192, unipunctus male and female thorax. Scale = 2 mm (189 redrawn from Barnard, 1980).

Figs. 193–201. Polymorphanisus male right fore- and hind wings. 193, astictus; 194, fuscus; 195, muluensis; 196, nigricornis; 197, ocularis; 198, quadripunctatus; 199, semperi; 200, scutellatus; 201, unipunctus. Scale = 5 mm (199 redrawn from Barnard, 1980).

Figs. 226–234. Polymorphanisus segment X of male genitalia. 226, astictus; 227, fuscus; 228, muluensis; 229, nigricornis; 230, ocularis; 231, quadripunctatus; 232, semperi; 233, scutellatus; 234, unipunctus. Scale = 0.5 mm (232 redrawn from Barnard, 1980).