Paraulacoryssus puertoricensis (Mutchler)

Shpeley, Danny, Hunting, Wesley & Ball, George E., 2017, A taxonomic review of the Selenophori group (Coleoptera, Carabidae, Harpalini) in the West Indies, with descriptions of new species and notes about classification and biogeography, ZooKeys 690, pp. 1-195 : 12-14

Paraulacoryssus puertoricensis (Mutchler) Figs 1B, 2 D–F, 3B, 4

Selenophorus puertoricensis Mutchler, 1934: 5. HOLOTYPE male: Desengano, Puerto Rico, December 1, W.T.M. Forbes (AMNH). PARATYPE female: Manidos, Puerto Rico, March 17, W.M. Wheeler (AMNH).- Darlington 1934: 104.- Blackwelder 1944: 50.- Erwin and Sims 1984: 440.- Ball 1992: 84, 85.- Lorenz 1998: 356.- Lorenz 2005: 377.

Type locality.

Desengano, Lajas Municipality, Puerto Rico.

Diagnosis.

This species is readily separated from all other West Indian selenophorine species by the reduced metepisternum, which has the anterior and lateral margins nearly equal in length.

Descriptive notes.

Data for SBL in Table 1. Habitus as in Fig. 1B. Labrum with anterior margin shallowly convex and clypeus with anterior margin shallowly concave. Antennae and mouthparts rufo-testaceous to nearly brunneous, with antennomere 1 paler than remainder of antenna. Legs rufo-brunneous to dark brunneous. Dorsal and ventral surfaces rufo-brunneous to brunneo-piceous. Elytra and ventral surface with faint iridescence. Head, pronotum and elytra shiny, without microlines visible at 100 ×. Pronotum with posteriolateral angles rounded; posteriolateral impressions and laterally near the bead finely punctate. Elytral striae impunctate, except the standard setigerous punctures in striae 2, 5 and 7. Intervals with fine micro-punctures. Both males and females with two terminal setae near the posterior margin on sternum VII.

Male genitalia. Fig. 2 D–F. Apical portion of phallic median lobe triangular, symmetrically rounded in dorsal/ventral aspect; endophallus with three fields of short spines, best seen in left lateral aspect; well sclerotized, sharply pointed lamina present, short, triangular in form, rounded on right, concave on the left.

Ovipositor and female reproductive tract. Fig. 3B. Gonocoxite 2 moderately thick, nearly straight. Bursa copulatrix short; small kidney-shaped spermathecal basal sclerite (sbs) and long, inflated spermatheca (sp) originating near base of common oviduct, terminated with one or two sausage like extensions; spermathecal gland duct originating well above base of spermatheca 2. Spermathecal gland duct long, bulbous swelling of duct basad triramous gland (spg).

Geographical distribution.

Fig. 4. This species is only known from the Greater Antillean island of Puerto Rico.

Chorological affinities and relationships.

See above for treatment of the genus Paraulacoryssus .

Material examined.

In addition to type material, we have seen a total of 8 specimens (2 males, 6 females). See Appendix for details.