Belisana protumida, Yao, Zhiyuan, Li, Shuqiang & Jäger, Peter, 2014

Yao, Zhiyuan, Li, Shuqiang & Jäger, Peter, 2014, Four new species of pholcine spiders (Araneae: Pholcidae) from Southeast Asia, Zootaxa 3793 (3), pp. 331-349 : 332-336

Belisana protumida spec. nov.

Figs 1–16

Type material. MALAYSIA: Peninsular Malaysia: holotype male, Taman Negara, 4°42′N, 102°28′E, shrubs, date and collector unknown ( SMF). Paratypes: 1 male, 1 female, same data as holotype ( SMF).

Etymology. The specific name is derived from the Latin protumidus (meaning protuberant), in reference to the small slender apophysis on the pedipalpal trochanter; adjective.

Diagnosis. Belisana protumida spec. nov. resembles B. martensi Yao & Li, 2013 (see Yao & Li 2013, figs 16–18, 41) in the shape of the membranous distal apophyses of the procursus, but can be distinguished by the absence of brown lateral margins on the dorsal shield of prosoma ( Figs 9–10 ); by the angular proximo-lateral apophyses and the relatively short distance between the tips of the distal apophyses on the male chelicerae ( Figs 8 , 14 ; proximo-lateral apophyses thumb-shaped and inter-distance between distal apophyses larger in B. martensi ); by the presence of a small slender apophysis on the pedipalpal trochanter ( Figs 1–2 , 12–13 ; apophyses absent in B. martensi ); and by the absence of stick-shaped apophyses on the procursus ( Figs 1–4 , 12–13 ; stick-shaped apophysis present in B. martensi ). This species also resembles B. bohorok Huber, 2005 (see Huber 2005a, figs 395–398, 403–410) and B. kaharian Huber, 2005 (see Huber 2005a, figs 399–402, 411–416) in having similarly shaped male chelicerae and narrow vulval pore plates, but can be distinguished by the absence of sclerites on the distal procursus ( Figs 1–4 , 12–13 ; flat sclerite present in B. bohorok and B. kaharian ).

Description. Male (holotype): Total length 1.28 (1.36 with clypeus), prosoma 0.53 long, 0.58 wide, opisthosoma 0.75 long, 0.45 wide. Leg I missing, leg II: – (2.28 + 0.22 + 1.97 + 2.97 + –), leg III: 5.11 (1.50 + 0.19 + 1.17 + 1.70 + 0.55), leg IV: 7.66 (2.25 + 0.21 + 1.88 + 2.63 + 0.69). Habitus as in Fig. 9 . Dorsal shield of prosoma and sternum yellowish, without marks. Legs yellowish, without darker rings. Opisthosoma yellowish, without spots. Distance PME-PME 0.16, diameter PME 0.08, distance PME-ALE 0.02, AME absent. Ocular area not elevated. Thoracic furrow absent. Sternum about as wide as long (0.43). Chelicerae as in Figs 8 , 14 , with a pair of angular proximo-lateral apophyses and a pair of cone-shaped distal apophyses (distance between tips of distal apophyses: 0.03). Pedipalp as in Figs 1–2 , 12–13 ; trochanter with a short retro-ventral apophysis provided with a small slender apophysis (arrow in Figs 1–2 , 12–13 ); femur with a dorsal apophysis; procursus simple proximally but complex distally, with a bent spine prolaterally and a membranous flap retrolaterally; bulb with a hooked apophysis and a simple embolus. Legs II, III and IV with short vertical hairs on metatarsi, without spines and curved hairs.

Variation: Total length in another male: 1.50 (1.61 with clypeus); leg I missing.

Female: Similar to male, habitus as in Figs 10–11 . Total length 1.52 (1.58 with clypeus), prosoma 0.55 long, 0.61 wide, opisthosoma 0.97 long, 0.47 wide; tibia I: 3.44; tibia I L/d: 55. Distance PME-PME 0.13, diameter PME 0.08, distance PME-ALE 0.01, AME absent. Epigyne ( Figs 5 , 15 ) simple and flat externally, with a pair of pockets 0.02 apart. Vulva ( Figs 6 , 16 ) with a ridge-shaped arch anteriorly and a pair of long, narrow pore plates. Retrolateral trichobothrium of tibia I at 11% of length; tarsus I missing.

Distribution. Known only from the type locality ( Fig. 57 ).

Figures

FIGURE 57. Known distribution records of four new pholcine species from Southeast Asia. 1, Belisana protumida spec. nov.; 2, Khorata bayeri spec. nov.; 3, Pholcus schawalleri spec. nov.; 4, Uthina khaosokensis spec. nov.

FIGURES 1 – 4. Belisana protumida spec. nov., holotype male. 1 – 2 Pedipalp (1 prolateral view, 2 retrolateral view; arrow points at a slender apophysis on pedipalpal trochanter); 3 – 4 Distal part of procursus (3 prolateral view, 4 retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = membranous flap, pr = procursus.

FIGURES 5 – 11. Belisana protumida spec. nov., holotype male (7, 9), paratype male (8), and paratype female (5 – 6, 10 – 11). 5 Epigyne, ventral view; 6 Vulva, dorsal view; 7 Bulb, prolateral view; 8 Chelicerae, frontal view; 9 – 11 Habitus (9 – 10 dorsal view, 11 ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, pa = proximo-lateral apophysis, pp = pore plate.

FIGURES 12 – 16. Belisana protumida spec. nov., holotype male (12 – 13), paratype male (14), and paratype female (15 – 16). 12 – 13 Pedipalp (12 prolateral view, 13 retrolateral view; arrow points at a slender apophysis on pedipalpal trochanter); 14 Chelicerae, frontal view; 15 Epigyne, ventral view; 16 Vulva, dorsal view. b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = epigynal pocket, f = membranous flap, pa = proximo-lateral apophysis, pp = pore plate, pr = procursus.