Amphineurus (Rhamphoneurus) stigmaticus, Santos & Santos & Ribeiro, 2022

Santos, Daubian, Santos, Rodrigo dos Reis & Ribeiro, Guilherme Cunha, 2022, Systematics of Amphineurus (Rhamphoneurus Alexander) (Diptera: Tipuloidea: Limoniidae), Arthropod Systematics & amp; Phylogeny 80, pp. 439-494 : 439

3.3.24. Amphineurus (Rhamphoneurus) stigmaticus sp. nov.

Material examined.

Holotype: ♂, Chile, Llanquihue, Hornohuinca [41°25′S 72°37′W], XII-1968, Peña (USNM)*. Paratypes: 1 ♀ and 1 [sex unknown], Chile, Arauco, Nahuelbuta [37°46′S 72°59′W], 5/ 10-II-1953, Peña (USNM) ; 1 ♂, Chile, Chiloé Is., Chepu [42°2′S 74°1′W], 4/ 6-IV-1968, Peña (USNM)*. - Additional material: CHILE. 2 ♂, Pucotrihue [40°32′S 73°42′W], 60 m, 10-III-1955, Peña (USNM) ; 1 male and 1 ♀, Chiloé Is. Ancud [41°52′S 73°48′W], 17-XII-1926, Peña (USNM); 1 ♂, Chiloé Is., Aulen [42°2′S 74°1′W], 8-II-1952, L.P. Guzman (USNM); 1 ♂ and 1 [sex unknown], Pucotrihue [40°32′S 73°42′W], 60 m, 1/ 13-III-1955, Peña (USNM); 1 [sex unknown], Llanquihue, Hornohuinca [41°25′S 72°37′W], South of Volcan Calbuco , 30/ 31-III-1968, Peña (USNM); 1 [sex unknown], Chiloé Is., Chepu [42°2′S 74°1′W], 1/ 3-II-1952, Peña (USNM) ; 1 male and 1 [sex unknown], Arauco, Nahuelbuta [37°46′S 72°59′W], 5/ 10-II-1953, Peña (USNM) ; 1 ♀, Casa Pangue [41°03′S 71°51′W], 4-XII-1926, R. C. Shannon (USNM); 1 [sex unknown], Chiloé Is., Chaiten [42°58′S 72°32′W], 5/ 8-II-1954, Peña (USNM); 1 [sex unknown], Chiloé Is., Aulen [42°2′S 74°1′W], 4-II-1952, Peña (USNM); 1 ♀ and 1 [sex unknown], Chiloé Is., Chepu [42°2′S 74°1′W], 4/ 6-IV-1968, Peña (USNM); 1 ♂, Arauco, Nahuelbuta, Caramavida [37°40′S 73°20′W], 1000 m, 5/ 10-II-1953, Peña (USNM) *; 1 ♂, Chiloé Is., Chaiten [42°58′S 72°32′W], 5/ 8-II-1954, Peña (USNM)*; 1 ♂, Chiloé Is., Chepu [42°2′S 74°1′W], 1/ 3-II-1951, Peña (USNM)*.

Etymology.

The word Amphineurus stigmaticus is Latin for “marked”. The term refers to the darkened mark near the tip of the wing of this new species.

Diagnosis.

This species is distinguished by the darkened distal quarter of the wings and basal deflection of M1 shorter than M1+2. Furthermore, the species is characterized by the male terminalia with setae on the medial branch of clasper of gonostylus and small marking on sheath of aedeagus.

Description.

Wing length 7.48 mm, width 2.57 mm. - Coloration: General coloration dark brown. Rostrum and palpus dark brown. First segment of the antenna yellow, remainding segments dark yellow. Head dark brown. Eye gray. Thorax dark brown, pleura brown. Scutum with three brownish black stripes. Halter pale. Coxae and legs brownish yellow. Abdomen dark brown. - Head (dorsal view Fig. 29B , ventral view Fig. 29A ): Rostrum bifurcated with short appendices; first palpal segment shorter than terminal segment; palpus with several setae; scape thicker than pedicel; terminal flagellomere as long as penultimate. - Thorax (lateral view Fig. 29E , dorsal view Fig. 29F ): Scutum with two rows of short setae; anatergite shorter than katatergite. Halter with dilated knob. Wing (Fig. 29D ) with several markings including faint marking on A1; markings highlighting m-cu, r-m, part of R5, and R2+3; marking on Rs, darkened distal quarter of wing and strong markings on R2 and at tip of R1; veins near fork of bM (Fig. 29C ) faded; M1+2 longer than basal deflection of M1; R2+3+4 shorter than half-length of R2+3. - Female terminalia (Fig. 29H ): Female tergite IX as wide and longer than tergite X; hypogynial valve slender; cercus bent at right angle, expanded in base; cercus shorter than hypogynial valve. - Male terminalia (Fig. 29G ): Male tergite IX with shallow median U-shaped notch. Dorsal branch of gonocoxite longer than half-length of ventral branch, well separated from ventral branch. Lobe of the gonostylus rounded; lobule longer than wide, shorter than half-length of lobe of gonostylus, narrower than stem. Clasper of gonostylus with differently shaped branches: lateral branch straightened; medial branch curved, round club shaped, with concentration of setae. Mesal lobes of gonocoxite symmetrical; both rounded with bump on side. Sheath of aedeagus angulated, darkened only near tip.

Remarks.

Specimens of this new species were previously identified by C.P. Alexander as A. (R.) nothofagetorum Alexander. This species resembles A. (R.) caleuchus sp. nov. but differs mainly in marking in the sheath of aedeagus, length of R2+3+4, shape of lobe of gonostylus, cercus and mediotergite.

Figures

Figure 29. Amphineurus (Rhamphoneurus) stigamticus sp. nov. A head (ventral view); B head (dorsal view); C detail of wing; D wing; E thorax (lateral view); F thorax (dorsal view); G male tergite IX (dorsal view) and remainder of male terminalia (lateral view); H female terminalia (dorsal view). - Abbreviations: aed sh, sheath of aedeagus; anatg, anatergite; anepm, anepimeron; anepst, anepisternum; aprn, anteropronotum; cer, cercus; cgonst, clasper of gonostylus; cx, coxa; goncx, gonocoxite; hlt, halter; hyp val, hypogynial valve; k, knob of halter; kepm, katepimeron; kepst, katepisternum; ktg, katatergite; l ms, left mesal lobe of gonocoxite; lgonst; lobe of gonostylus; mr, meron; ms lobe, mesal lobe of gonocoxite; mtanepst, metanepisternum; mtepm, metepimeron; mtg, mediotergite; mtkepst, metakatepisternum; mtn, metanotum; p, posterior basalare; patg, paratergite; pprn, postpronotum; presct, prescutum; r ms, right mesal lobe of gonocoxite; sct, scutum; sctl, scutellum; st, sternite; tg, tergite; t 9, male tergite IX.