Namadytes cimbebasiensis Hesse, 1972

Dikow, Torsten & Leon, Stephanie, 2014, Review of the genus Namadytes Hesse, 1969 (Insecta: Diptera: Mydidae: Syllegomydinae), Biodiversity Data Journal 2, pp. 1071-1071 : 1071

Namadytes cimbebasiensis Hesse, 1972

Namadytes cimbebasiensis Hesse, 1972: 143.

Materials

Type status: Holotype. Occurrence: catalogNumber: SAM-DIP-A007146 ; recordedBy: H. Brown; sex: 1 male; lifeStage: Adult; otherCatalogNumbers: AAM-000450; Taxon: scientificNameID: urn:lsid:zoobank.org:act:5AA365A2-2479-4959-BF33-45C64E746A1F; scientificName: Namadytescimbebasiensis Hesse, 1972; family: Mydidae; genus: Namadytes; specificEpithet: cimbebasiensis; scientificNameAuthorship: Hesse, 1972; Location: country: Namibia; stateProvince: Hardap; county: Maltahöhe; locality: Excelsior No. 127 ; verbatimCoordinates: 25°24'00''S 016°12'00''E; decimalLatitude: -25.4; decimalLongitude: 16.2; Identification: identifiedBy: A. Hesse; dateIdentified: 1972; Event: eventDate: 1969-05-07; Record Level: institutionCode: SAMC; collectionCode: Insects; basisOfRecord: PreservedSpecimen

Type status: Paratype. Occurrence: catalogNumber: SAM-DIP-A007146 ; recordedBy: H. Brown; sex: 1 female; lifeStage: Adult; otherCatalogNumbers: AAM-000451; Taxon: scientificNameID: urn:lsid:zoobank.org:act:5AA365A2-2479-4959-BF33-45C64E746A1F; scientificName: Namadytescimbebasiensis Hesse, 1972; family: Mydidae; genus: Namadytes; specificEpithet: cimbebasiensis; scientificNameAuthorship: Hesse, 1972; Location: country: Namibia; stateProvince: Hardap; county: Maltahöhe; locality: Excelsior No. 127 ; verbatimCoordinates: 25°24'00''S 016°12'00''E; decimalLatitude: -25.4; decimalLongitude: 16.2; Identification: identifiedBy: A. Hesse; dateIdentified: 1972; Event: eventDate: 1969-05-07; Record Level: institutionCode: SAMC; collectionCode: Insects; basisOfRecord: PreservedSpecimen

Type status: Paratype. Occurrence: catalogNumber: SAM-DIP-A007146 ; recordedBy: H. Brown; sex: 1 male; lifeStage: Adult; otherCatalogNumbers: AAM-000452; Taxon: scientificNameID: urn:lsid:zoobank.org:act:5AA365A2-2479-4959-BF33-45C64E746A1F; scientificName: Namadytescimbebasiensis Hesse, 1972; family: Mydidae; genus: Namadytes; specificEpithet: cimbebasiensis; scientificNameAuthorship: Hesse, 1972; Location: country: Namibia; stateProvince: Hardap; county: Maltahöhe; locality: Excelsior No. 127 ; verbatimCoordinates: 25°24'00''S 016°12'00''E; decimalLatitude: -25.4; decimalLongitude: 16.2; Identification: identifiedBy: A. Hesse; dateIdentified: 1972; Event: eventDate: 1969-05-07; Record Level: institutionCode: SAMC; collectionCode: Insects; basisOfRecord: PreservedSpecimen

Type status: Paratype. Occurrence: catalogNumber: SAM-DIP-A007146 ; recordedBy: H. Brown; sex: 1 male; lifeStage: Adult; otherCatalogNumbers: AAM-000453; Taxon: scientificNameID: urn:lsid:zoobank.org:act:5AA365A2-2479-4959-BF33-45C64E746A1F; scientificName: Namadytescimbebasiensis Hesse, 1972; family: Mydidae; genus: Namadytes; specificEpithet: cimbebasiensis; scientificNameAuthorship: Hesse, 1972; Location: country: Namibia; stateProvince: Hardap; county: Maltahöhe; locality: Excelsior No. 127 ; verbatimCoordinates: 25°24'00''S 016°12'00''E; decimalLatitude: -25.4; decimalLongitude: 16.2; Identification: identifiedBy: A. Hesse; dateIdentified: 1972; Event: eventDate: 1969-05-07; Record Level: institutionCode: SAMC; collectionCode: Insects; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: NMNW-H7809 ; sex: 1 female; lifeStage: Adult; otherCatalogNumbers: AAM-003014; Taxon: scientificNameID: urn:lsid:zoobank.org:act:5AA365A2-2479-4959-BF33-45C64E746A1F; scientificName: Namadytescimbebasiensis Hesse, 1972; family: Mydidae; genus: Namadytes; specificEpithet: cimbebasiensis; scientificNameAuthorship: Hesse, 1972; Location: country: Namibia; stateProvince: Karas; county: Keetmanshoop; locality: Swartbaas West No. 276 ; verbatimCoordinates: 27°00'16''S 019°41'08''E; decimalLatitude: -27.00444; decimalLongitude: 19.68556; Identification: identifiedBy: T. Dikow S. Leon; dateIdentified: 2012; Event: eventDate: 1972-04-19-1972-04-22; Record Level: institutionCode: NMNW; collectionCode: Insects; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: NMNW-H7808 ; sex: 1 male; lifeStage: Adult; otherCatalogNumbers: AAM-003013; Taxon: scientificNameID: urn:lsid:zoobank.org:act:5AA365A2-2479-4959-BF33-45C64E746A1F; scientificName: Namadytescimbebasiensis Hesse, 1972; family: Mydidae; genus: Namadytes; specificEpithet: cimbebasiensis; scientificNameAuthorship: Hesse, 1972; Location: country: Namibia; stateProvince: Karas; county: Keetmanshoop; locality: Swartbaas West No. 276 ; verbatimCoordinates: 27°00'16''S 019°41'08''E; decimalLatitude: -27.00444; decimalLongitude: 19.68556; Identification: identifiedBy: T. Dikow S. Leon; dateIdentified: 2012; Event: eventDate: 1972-04-19-1972-04-22; Record Level: institutionCode: NMNW; collectionCode: Insects; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: AAM-003000 ; recordedBy: M. Schwarz; sex: 1 female; lifeStage: Adult; otherCatalogNumbers: AAM-003000; Taxon: scientificNameID: urn:lsid:zoobank.org:act:5AA365A2-2479-4959-BF33-45C64E746A1F; scientificName: Namadytescimbebasiensis Hesse, 1972; family: Mydidae; genus: Namadytes; specificEpithet: cimbebasiensis; scientificNameAuthorship: Hesse, 1972; Location: country: South Africa; stateProvince: Northern Cape; locality: Van Zylsrus, 90 km W ; verbatimCoordinates: 27°04'47''S 021°17'08''E; decimalLatitude: -27.07972; decimalLongitude: 21.28556; Identification: identifiedBy: T. Dikow S. Leon; dateIdentified: 2012; Event: eventDate: 1990-03-26; Record Level: institutionCode: CSCA; collectionCode: Insects; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: AAM-003021 ; recordedBy: F. and S. Gess; sex: 1 male; lifeStage: Adult; otherCatalogNumbers: AAM-003021; Taxon: scientificNameID: urn:lsid:zoobank.org:act:5AA365A2-2479-4959-BF33-45C64E746A1F; scientificName: Namadytescimbebasiensis Hesse, 1972; family: Mydidae; genus: Namadytes; specificEpithet: cimbebasiensis; scientificNameAuthorship: Hesse, 1972; Location: country: South Africa; stateProvince: Northern Cape; locality: 58 km N on R360 Upington-Kgalagadi ; verbatimCoordinates: 27°59'22''S 020°59'51''E; decimalLatitude: -27.98944; decimalLongitude: 20.9975; Identification: identifiedBy: T. Dikow S. Leon; dateIdentified: 2012; Event: eventDate: 2000-04-06; Record Level: institutionCode: AMGS; collectionCode: Insects; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: NMSA-DIP-66615 ; recordedBy: J. Londt; sex: 1 male; lifeStage: Adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:5AA365A2-2479-4959-BF33-45C64E746A1F; scientificName: Namadytescimbebasiensis Hesse, 1972; family: Mydidae; genus: Namadytes; specificEpithet: cimbebasiensis; scientificNameAuthorship: Hesse, 1972; Location: country: South Africa; stateProvince: Northern Cape; locality: Kgalagadi Transfontier Park, Kalahari Tented Camp ; verbatimElevation: 947 m; verbatimCoordinates: 25°47'08''S 020°01'01''E; decimalLatitude: -25.78556; decimalLongitude: 20.01694; Identification: identifiedBy: T. Dikow; dateIdentified: 2013; Event: eventDate: 2012-04-11; habitat: dune scrub; Record Level: institutionCode: NMSA; collectionCode: Insects; basisOfRecord: PreservedSpecimen

Type status: Other material. Occurrence: catalogNumber: NMSA-DIP-67240 ; recordedBy: J. and A. Londt; sex: 1 male; lifeStage: Adult; Taxon: scientificNameID: urn:lsid:zoobank.org:act:5AA365A2-2479-4959-BF33-45C64E746A1F; scientificName: Namadytescimbebasiensis Hesse, 1972; family: Mydidae; genus: Namadytes; specificEpithet: cimbebasiensis; scientificNameAuthorship: Hesse, 1972; Location: country: South Africa; stateProvince: Northern Cape; locality: Kgalagadi Transfontier Park, Twee Rivieren ; verbatimElevation: 864 m; verbatimCoordinates: 26°28'27''S 020°36'46''E; decimalLatitude: -26.47417; decimalLongitude: 20.61278; Identification: identifiedBy: T. Dikow; dateIdentified: 2013; Event: eventDate: 2012-04-08-2012-04-14; habitat: Acacia savanna; Record Level: institutionCode: NMSA; collectionCode: Insects; basisOfRecord: PreservedSpecimen

Description

Male: Fig. 5a, b.

Head: brown, in general grey pubescent; width distinctly greater than thorax, interocular distance on vertex larger than at ventral eye margin, vertex between compound eyes ± horizontally straight, medially only slightly below dorsal eye margin, parafacial area about as wide as ½ the width of central facial gibbosity; facial gibbosity distinct, well-developed and discernible in lateral view; mystax yellow, covering entire facial gibbosity; frons not elevated, predominantly apubescent; vertex entirely grey pubescent; postgena lightly grey pubescent; setation: vertex yellow, frons white or yellow, ocp setae white, pocl macrosetae absent; ocellar triangle apubescent; proboscis light brown, very short, vestigial, knob-like; labellum small, as wide as prementum, as long as prementum, unsclerotized laterally; maxillary palpus cylindrical, yellow, minute.

Antenna: brown, scape and pedicel white setose dorsally and ventrally; postpedicel cylindrical in proximal ½, symmetrically bulbous in distal ½, ≥ 4.0 times as long as combined length of scape and pedicel, asetose; apical seta-like sensory element situated apically in cavity on postpedicel.

Thorax: brown, lightly grey pubescent; scutum uniformly black or uniformly brown, surface entirely smooth, lightly grey pubescent, broad sublateral stripes apubescent, scutal setation comprised of scattered long white to yellow setae; dc setae pre- and postsuturally white or yellow, acr setae absent, lateral scutal setae white or yellow, npl setae 0, spal setae 0, pal setae 0; antepronotum dorso-medially with V-shaped indentation; postpronotal lobe light brown, grey pubescent; proepisternum, lateral postpronotum, and postpronotal lobe long white setose; scutellum grey pubescent, asetose, apical scutellar setae absent; mesopostnotum, anatergite, and katatergite partly grey pubescent, mesopostnotum apubescent, mesopostnotum asetose, anatergite asetose, katatergite long white setose; katatergite ± flat; anterior anepisternum white setose, supero-posterior anepisternum long white setose; posterior anepimeron asetose, katepimeron asetose; metanepisternum grey pubescent, asetose, metepimeron ± flat, yellow, grey pubescent, long white setose; infra-halter sclerite asetose or white setose.

Leg: light brown, setation yellow; pro, mes, and met coxa lightly white pubescent, short yellow setose; met trochanter setose medially; femur brown or light brown, met femur ± cylindrical only slightly wider than pro and mes femur, in distal ½ macrosetose, 1 antero-ventral and 1 postero-ventral row of macrosetae, postero-ventrally regular setose only; pro, mes, and met tibia straight, met tibia cylindrical, ventral keel absent, latero-posteriorly regular setose only; pro and mes tarsomere 1 longer than tarsomere 2, but less than combined length of tarsomeres 2-3, met tarsomere 1 as long as combined length of tarsomeres 2-4; pulvillus well-developed, as long as well-developed claw, and as wide as base of claw; empodium absent.

Wing: length = 6.6-8.1 mm; hyaline throughout, veins light brown, microtrichia absent; cells r1, r4, r5, m3, + cup closed; C terminates at junction with R1; R4 terminates in R1; R5 terminates in R1; stump vein (R3) at base of R4 present, short not reaching R2; R4 and R5 widest apart medially; r-m distinct, R4+5 and M1 apart, connected by crossvein; M1 straight at r-m (not curving anteriorly), M1 (or M1+M2) terminates in R1; CuA1 and CuA2 split proximally to m-cu (cell m3 narrow proximally); M3+CuA1 do not terminate together in C; A1 undulating, cell a1 wide, A1 and wing margin further apart proximally than distally; alula well-developed; halter light yellow.

Abdomen: yellow to brown; setation comprised of dense white setae, surface entirely smooth; T1 brown, T2 brown anteriorly and postero-medially, otherwise yellow, T3-7 brown and yellow posteriorly; T1 and anterior ½ of T2 long white setose, remaining T short white setose; T predominantly apubescent; S1-7 light brown; S1 asetose, S2-7 sparsely white setose; S predominantly apubescent; T2-4 parallel-sided and not constricted waist-like; bullae on T2 black, transversely elongate, surface entirely smooth, T2 surface anterior to bullae smooth.

♂ terminalia: Fig. 2a, b, c.

Female: Fig. 5c, d.

Head: brown, facial gibbosity light brown; mystax white or yellow, sparsely covering entire facial gibbosity; setation: vertex white or yellow, pocl macrosetae light brown; maxillary palpus light brown.

Thorax: light brown, predominantly grey pubescent; scutum uniformly brown, predominantly brown pubescent, narrow sublateral stripes (wider anteriorly) and lateral and posterior margins grey pubescent, scutal setation comprised of scattered short yellow setae; proepisternum, lateral postpronotum, and postpronotal lobe short white setose; mesopostnotum, anatergite, and katatergite lightly grey pubescent, katatergite short white setose; anterior anepisternum white to yellow setose, supero-posterior anepisternum short white to yellow setose; metepimeron light brown; infra-halter sclerite asetose.

Leg: femur brown; pulvillus reduced, half length of well-developed claw.

Wing: length = 9.6-12.7 mm.

Abdomen: setation comprised of sparsely scattered short yellow setae; T1-7 brown, T2-6 light brown medially; T1-7 sparsely yellow setose; S1-7 brown; S1-7 sparsely short yellow setose; bullae on T2 oval.

♀ genitalia (Fig. 1): 8-10 acanthophorite spines per plate.

Diagnosis

This rather small species (wing length in males 6.6-8.1 mm and in females 9.6-12.7 mm) is distinguished from congeners by the entirely grey pubescent vertex, the short postpedicel (only about 4 times as long as combined length of scape and pedicel), the grey pubescent scutellum, and the few white setae on the infra-halter sclerite, which are absent in some females.

Distribution

Namibia (Hardap, Karas) and South Africa (Northern Cape) (Fig. 6).

Biology

Habitat

Namadytes cimbebasiensis has recently been collected on dune scrub and in Acacia savanna in the Kgalagadi Transfrontier Park of the Kalahari Desert by J.G.H. Londt.

Discussion

Namadytes cimbebasiensis is very distinct and the smallest species of Namadytes . All specimens, originating from only five collecting events, are either greasy or are not in the best condition so that in particular the pubescence patterns might differ in freshly mounted material.

Type locality

Namibia: Hardab: Excelsior No. 127 (25°24'00''S, 016°12'00''E) (Fig. 6).

Biodiversity hotspot

Not known to occur in any of the southern African biodiversity hotspots (Cape Floristic Region, Maputaland-Pondoland-Albany, or Succulent Karoo) (Fig. 6).