Colura ari (Steph.) Steph.

Sarimi, Muhammad Syafiq, Pócs, Tamás & Lee, Gaik Ee, 2021, Data to the Malaysian liverwort flora, III: New Lejeuneaceae records from Sabah and Peninsular Malaysia, Cryptogamie, Bryologie 20 (19), pp. 249-267 : 254

Colura ari (Steph.) Steph.

( Figs 10 ; 11C, D )

Species Hepaticarum 5: 936 (1916).

SPECIMEN EXAMINED. — Malaysia. Terengganu: Mt. Tebu Forest Reserve, 15 km S of Jerteh town. Primary lowland dipterocarp forest with small waterfalls and slow stream. Along the forest trail at the Lata Belatan Recreational Forest and waterfall, situated at the base of Mt. Tebu at 100 m alt., on living leaves, 16.IV.2019, G.E. Lee et al. 19018, 19019, 19020, 19021.

REMARKS

Distinctive features of Colura ari include the strongly serrate leaf margin, the position of lobule and sac more or less parallel to the stem, truncate and apiculate sac (apex usually pointing downward), and leaf cells with large trigones and conspicuous intermediate thickenings (2 per adjacent trigones in basal leaf cells). A common and widespread Indopacific epiphyllous species. New to Peninsular Malaysia (Terengganu). In Malaysia, only known from Sabah ( Mizutani 1966, 1970).

Figures

FIG. 10. — Colura ari (Steph.) Steph.: A, part of plant in ventral view; B, C, leaves; D, underleaf; E, marginal leaf lobe cells; F, valve; G, median leaf lobe cells; H, apex of lobule sac; I, perianth. From G.E. Lee et al. 19020. Scale bars: A-C, I, 0.5 mm; D, 0.2 mm; E, G, 100 µm; F, H, 50 µm.

FIG. 11. — Colura inuii Horik.: A, habit; B, oil bodies. From G.E.Lee 21010. Colura ari (Steph.) Steph.: C, habit; D, oil bodies. From G.E. Lee et al. 19020. Radula grandilobula Promma & Chantanaorr.: E, habit; F, oil bodies. From G.E. Lee 21003. Scale bars: A, C, E, 2 mm; B, D, 50 µm; F, 25 µm.