Cololejeunea appressa (A.Evans) Benedix

Sarimi, Muhammad Syafiq, Pócs, Tamás & Lee, Gaik Ee, 2021, Data to the Malaysian liverwort flora, III: New Lejeuneaceae records from Sabah and Peninsular Malaysia, Cryptogamie, Bryologie 20 (19), pp. 249-267 : 252

Cololejeunea appressa (A.Evans) Benedix

( Fig. 4 )

Feddes Repertorium Specierum Novarum Regni Vegetabilis. Beiheft

134: 31 (1953).

SPECIMEN EXAMINED. — Malaysia. Terengganu: Mt. Tebu Forest Reserve, 15 km S of Jerteh town. Primary lowland dipterocarp forest with small waterfalls and slow stream. Along the forest trail at the Lata Belatan Recreational Forest and waterfall, situated at the base of Mt. Tebu at 100 m alt., on living leaves, 16.IV.2019, G.E. Lee et al. 19011.

REMARKS

Cololejeunea appressa is a member of subg. Taeniolejeunea and is distinguished from other members in this subgenus by the single row of 3-4 ocelli, the unicellular first lobule tooth and first and second lobule teeth clearly visible, not overlapping each other. A pantropical species, preferably growing on barks or on living leaves at low elevations, reported in Asia from China, Indonesia (Java, Sumatra), Indonesian New Guinea, Japan, and recently from Laos ( Mizutani 1961; Piippo 1990; Pócs & Kovács 2019; Söderström et al. 2020). New to Peninsular Malaysia (Terengganu). In Malaysia the species is only known from Mt. Kinabalu, Sabah ( Mizutani 1966). In view of its wide world distribution, however, this species is expected to occur in many lowland forest areas in Malaysia.

Figures

FIG. 4. — Cololejeunea appressa (A.Evans) Benedix:A, part of plant in ventral view; B, C, F, leaves;D, stem portion,leaf lobule and stylus (hyaline papilla shown in gray); E, marginal leaf lobe cells; G, median leaf lobe cells; H, basal leaf lobe cells and ocelli. From G.E. Lee et al. 19011. Scale bars: A, 0.2 mm; B, C, F, 0.1 mm; D, H, 50 µm; E, G, 25 µm.