Philonthus carbonarius (Gravenhorst)

Pietrykowska-Tudruj, Ewa & Staniec, Bernard, 2011, The morphology of the pupae of six species of Philonthina (Coleoptera, Staphylinidae, Staphylinini) with taxonomic remarks, Zootaxa 2865, pp. 53-67 : 58

Philonthus carbonarius (Gravenhorst)

( Figs 10–12 , 26–32 , 47–48A, 55, 66)

Body length: 4.29–4.95 mm (mean 4.7 mm), female 4.57–4.95 mm (mean 4.81 mm), male 4.29–4.73 mm (mean 4.60 mm); body width: 1.93–2.20 mm (mean 2.00 mm), female 2.00– 2.22 mm (mean 2.1 mm), male 1.91–2.10 mm (mean 2.00 mm); head width: 1.06–1.19 mm (mean 1.13 mm); pronotum width: 1.39–1.65 mm (mean 1.53 mm), 1.1 x as broad as long; body rather stocky and weakly sclerotized. Head about 1.2 x as long as wide ( Figs 10, 11 ). Antennae protruding middle of elytra length ( Fig. 11 ). Anterior margin of pronotum with 10–13 projections (females–5/5, 7/6, 6/6, 7/6, 5/6 projections, males–6/6, 6/6, 5/6, 6/6, 6/6, 6/6, 6/6 projections on sides in individuals). The wing length as in G. osseticus and P. atratus ( Figs 4, 5 , 7, 8, 10, 11 ). Each fore, middle, and hind tibiae with 1, 9, and 4 protuberances respectively. The length of hind legs as in P. a t r a t u s ( Figs 7, 8, 10, 11 ). Abdominal segments II–IV almost equal in width, than narrowed to the terminal segment of the body ( Fig. 12 ). Abdominal segments III–VIII each with a pair of setiform projections (6 pairs in total) on sides ( Figs 10–12 ). Segments III–VI more than four (4.3–4.5 x) as long as projections ( Figs 28, 31 ); projections on segments VII and VIII, 3– 4 x longer than on the other segments; projections on segment VII only about 1.5x shorter than segment, projection on segment VIII curved, slightly longer than segment ( Figs 26, 30 ). All abdominal projections on sides bearing cuticular processes ( Figs 27, 29, 32 ). Abdominal segments with microstructure as in Fig. 26 A. Female pupa with two well-developed ventral, elongated, straight prolongations (Fig. 47); sternite IX of male pupa as in Fig. 48; ventral and terminal prolongation with tiny cuticular processes (Figs 47A, 48A). The first pair of functional spiracles slightly narrowed to the apex, with strongly-marked reticulate microstructure ( Figs 11, 12 , 55 ); atrophied spiracles as in Fig. 66 .

Figures

FIGURES 7 – 12. Pupa of Philonthus atratus (7 – 9) and P. carbonarius (10 – 12). (7, 10) Ventral aspect, (8, 11) lateral aspect, (9, 12) dorsal aspect.

FIGURES 19 – 40. Pupa of Gabrius osseticus (19, 20), Philonthus atratus (21 – 25), P. carbonarius (26 – 32), P. debilis (33 – 35) and P. decorus (36 – 40). (19, 21, 26, 27, 36) Lateral part of segment VII with microstructure (19 A, 26 A) and spine (27); (20, 22, 33, 39, 40) lateral part of segment VIII with microstructure (39 A) and spine (40); (23, 24, 28, 29) lateral part of segment IV with spine (24, 29); (25, 31, 32, 34, 35, 37, 38) lateral part of segment V with microstructure (34 A) and spine (32, 35, 38).

FIGURES 1 – 6. Pupa of Bisnius fimentarius (1 – 3) and Gabrius osseticus (4 – 6). (1, 4) Ventral aspect, (2, 5) lateral aspect, (3, 6) dorsal aspect.

FIGURES 52 – 70. Pupa of Bisnius fimentarius (52, 61, 62), Gabrius osseticus (53, 58, 63, 64), Philonthus atratus (54, 59, 65), P. carbonarius (55, 66), P. debilis (56, 60, 68 – 70) and P. d e c o r u s (57, 67). (52 – 60) The first pair of functional spiracles; (61 – 70) atrophied spiracles.