Omphale theana (Walker)

Hansson, Christer & Shevtsova, Ekaterina, 2012, Revision of the European species of Omphale Haliday (Hymenoptera, Chalcidoidea, Eulophidae), ZooKeys 232, pp. 1-157 : 39-42

Omphale theana (Walker) Figures 123-137485510539

Entedon theana Walker, 1839:81. Lectotype female in BMNH, examined.

Entedon ithonus Walker, 1839:82. Lectotype female in BMNH, examined. Synonymized by Graham (1963:254).

Derostenus (Derostenus) radialis Thomson, 1878:269. Lectotype female in LUZM, examined. Syn. n.

Omphale radialis (Thomson), Dalla Torre (1898).

Achrysocharella americana Girault, 1916:295. Holotype female in USNM, examined. Syn. n.

Omphale theana (Walker), Graham (1959).

Omphale theana (Walker), Graham (1963).

Omphale radialis (Thomson), Graham (1963).

Omphale theana (Walker), Hansson (1996b).

Omphale radialis (Thomson), Hansson (1996b).

Material.

Type material. Lectotype females of Entedon theana , type no. 5.2033, and Entedon ithonus , type no. 5.2034, both types in BMNH; lectotype female of Derostenus radialis , type no. 117:1 in LUZM; holotype female of Achrysocharella americana , type no 19586 in USNM. Additio-nal material. 334♀ 3♂: France 11♀ 1♂ (BMNH, RMNH), Germany 6♀ (RMNH), Hungary 10♀ (BMNH, CH), Netherlands 3♀ (RMNH), Russia 64♀ (BMNH, CH, LUZM), Sweden 175♀ (BMNH, CH, LUZM), United Kingdom 65♀ 2♂ (BMNH).

Diagnosis.

Female with 7th gastral tergite 1.3 –4.3× (average= 2.5, n= 156) as long as its basal width; interscrobal area, prosternum, prepectus and sides of scutellum frequently brownish-testaceous non metallic; coxae yellowish brown to yellowish white (Fig. 123), base with or without infuscation; forewing with 2-5 admarginal setae; female flagellomeres 2-4 ventrally with one set of setae attached subbasally and reaching beyond apex of flagellomere attached to (Fig. 131); male flagellomeres 1-4 with a single basal whorl of setae (Fig. 135), flagellomere 5 with scattered setae; forewing interference pattern, both sexes, with one colour (Fig. 129) - hue varies depending on size of specimen, which is different from the other species in this group that have different colours on basal and apical halves of the forewing. Very similar to Omphale acuminata , differs in the length of the 7th gastral tergite in female, which is longer in Omphale theana , see above "Morphometric analyses …”, and in having a different WIP - forewing with one colour in Omphale theana (Fig. 129), but with two large areas with different colours in Omphale acuminata (Fig. 77). Through the elongate female gaster also similar to Omphale salicis , from which it differs in having fewer admarginal setae in forewing (6-14 in Omphale salicis ), in having coxae predominantly non-metallic, and in WIPs - the unicoloured forewing WIP in Omphale theana separates it from the other species in this group.

Description.

Female. Length of body 1.2-2.3 mm. Antenna with scape yellowish brown with apical ⅓ and dorsal margin dark brown; pedicel and flagellum dark brown; pedicel + flagellum 2.1 × as long as distance between eyes; first flagellomere 1.0 × as long and 1.0 × as wide as second flagellomere (Fig. 131); flagellomeres 1-4 ventrally with one set of setae attached subbasally and reaching beyond apex of flagellomere attached to; clava 2-segmented. Face golden green to bluish green metallic (Fig. 126), strigose (Fig. 132); clypeus golden green with margin towards frons frequently brown, wrinkled to smooth, trapezoid to semicircular, 1.7 × as wide as high; gena not visible on type of Omphale theana but brown with golden tinges in non-type material; lower frons golden green to bluish green metallic, with weak reticulation; interscrobal area pale brown, yellowish brown, or dark brown metallic, smooth; antennal scrobes join frontal suture separately; frontal suture V-shaped; upper frons golden to bluish green metallic, with very weak reticulation; vertex brown with golden to golden green tinges, with very weak reticulation (Fig. 133). Occipital margin rounded (Fig. 133).

Mesoscutum golden red with green metallic tinges, golden green, or bluish green metallic (Fig. 124), with engraved reticulation (Fig. 130)), midlobe with two pairs (1-2 pairs in non-type material) of setae; notauli as indistinct impressions in posterior ½. Scutellum golden red, golden green, or bluish green metallic (Fig. 124), and frequently partly brown to yellowish brown - then usually with lateral parts non metallic, with engraved and weak reticulation (Fig. 130), 1.1 × as long as wide, with anterior margin smoothly curved forward. Axillae brown with golden tinges (Fig. 124). Dorsellum brown to yellowish brown (Fig. 124), convex and smooth (Fig. 130), 0.4 × as long as wide, and 1.0 × as long as length of median propodeum. Entire lateral mesosoma brown with metallic tinges (Fig. 123), to with propleuron, lateral pronotum, prepectus and mesepisternum yellowish brown; transepimeral sulcus curved forwards. Propodeum dark brown with metallic tinges (Fig. 124), smooth (Fig. 130); propodeal callus with two setae. Legs yellowish brown, with base of hind coxa brown (Fig. 123); midleg with first tarsal segment 0.4 × as long as tarsus. Forewing transparent, veins yellowish brown, setae dark brown (Fig. 128); speculum closed; admarginal setae 4 (2-5 in non-type material), arising from marginal vein; radial cell bare and long, 2.0 × as long as length of postmarginal vein; postmarginal vein 0.9 –1.0× as long as stigmal vein. Hind wing transparent, apex pointed (Fig. 128). Forewing WIP (Fig. 129) entirely in magenta.

Petiole brown. Gaster dark brown, to yellowish brown with posterior margin of tergites dark brown, and shiny, smooth, elongate and 2.8 × (2.0 –2.7× in non-type material) as long as length of mesosoma; 7th tergite 0.3 –0.5× as long as length of gaster.

Male. Length of body 1.2-1.3 mm. Features as in female except as follows. Antennal pedicel + flagellum 3.1 × as long as distance between eyes; flagellomeres 1-4 with verticillate setae and with setae reaching beyond apex of flagellomere attached to (Fig. 135); clava 1-segmented. Face metallic bluish green (Fig. 127), strigose (Fig. 136); clypeus metallic bluish green, smooth, 1.2 × as wide as high; gena golden green; lower frons golden green or golden red; upper frons golden or golden red; vertex golden red, with weak reticulation (Fig. 137).

Mesoscutum bluish green metallic (Fig. 125) or golden red, midlobe with one pair of setae (posterior pair) (Fig. 134). Scutellum golden green or golden red, 1.2 × as long as wide. Axillae golden. Dorsellum dark brown with golden tinges (Fig. 125); 0.7 × as long as length of median propodeum. Entire lateral mesosoma golden green or bluish green metallic. Propodeum golden red. Legs yellowish brown, hind coxa with base dark brown metallic; midleg with first tarsomere 0.3 × as long as length of tarsus. Forewing admarginal setae 3-5; postmarginal vein 0.9 × as long as stigmal vein.

Petiole dark brown. Gaster dark brown metallic, smooth, 1.1 × as long as length of mesosoma. Phallobase and aedeagus as in Fig. 485.

Host.

Unknown.

Distribution.

Czech Republic ( Bouček and Askew 1968), Faroes ( Kryger and Schmiedeknecht 1938), France ( Gijswijt 1976), Germany ( Gijswijt 1976), Hungary ( Erdös 1956), Netherlands ( Gijswijt 1976), Russia ( Yefremova 2002), Sweden ( Thomson 1878), United Kingdom ( Walker 1839); Canada ( Hansson 1996b), USA ( Girault 1916) (Fig. 510).