Pella Stephens, 1835

Hlaváč, Peter, Newton, Alfred F. & Maruyama, Munetoshi, 2011, World catalogue of the species of the tribe Lomechusini (Staphylinidae: Aleocharinae) 3075, Zootaxa 3075, pp. 1-151 : 68-71

Pella Stephens

Pella Stephens, 1833: 95 [= Stephens, 1835: 434]. Type species: Staphylinus limbatus Paykull , designated by Westwood, 1838: 20

Pella Stephens : Kistner, D. H. & M. S. Blum, 1971 (behavior); Kistner, 1972a: 150 ( Pella , redescription, moved to the generic level); Dvořák, 1980 (key to Czechoslovak species); Maruyama, 2006a (biosystematic revision of palearctic species, morphology, key); Klimaszewski et al. 2005: 716 (redescription, key to species from America north of Mexico)

Zyras (Lepla) Tottenham, 1939a: 226 . Type species: Aleochara lugens Gravenhorst , by original designation, synonymy in Maruyama, 2006a: 28

Myrmedonia (Myrmelia) Mulsant & Rey, 1873a: 152 [= Mulsant & Rey, 1873b: 54, 1874a: 6, 1874b: 86]. Type species: Myrmedonia excepta Mulsant & Rey , by monotypy, synonymy in Maruyama, 2006a: 27

Myrmedonia (Pellochromonia) Reitter, 1909: 43 . Type species: Myrmedonia ruficollis Grimm , by monotypy, synonymy in Maruyama, 2006a: 27

angustula Casey, 1894: 323 ( Myrmedonia, TL : Florida) Distribution: USA (Florida); new combination ex Zyras, Klimaszewski et al. 2005

barbara Fairmaire, 1863: 638 ( Myrmedonia, TL : Bone) Distribution: Algeria beijingorum Pace, 1998: 976 ( Zyras (Pella), TL : China, Beijing, Xiaolongmen, 1100-1500 m; HA: Lasius (Lasius) sp., L. (Chthonolasius) umbratus , L. (Dendrolasius) fuji , L. (D.) spathepus , L. (D.) capitatus ) Distribution: China (Beijing region), Japan (Hokkaidô, Honshû), far east Russia bohaci Dvořák, 1984: 195 ( Zyras (Pella), TL : UdSSR, Uzbekistan, Kyzyl-Kum, Šafirkan bei Buchara) Distribution: Kazakhstan, Turkmenistan, Uzbekistan

= almaatensis Pace, 2002a: 195 ( Zyras (Pella), TL : Kasachstan, Alma-Ata Gebiet, Pristan Dubinskaja) synonymy in Maruyama, 2006a: 139

brasiliana Wendeler, 1956: 270 ( Zyras (Pella), TL : Brasilien, Espirito Santo, Sta. Theresa) Distribution: Brazil caliginosa Casey, 1894: 323 ( Myrmedonia, TL : Catskill Mts., New York & Indiana) Distribution: USA (IN,NY,WV,WI,WA,FL,CA); new combination ex Zyras, Klimaszewski et al. 2005

carolinae Casey, 1911: 72 ( Myrmedonia, TL : Tryon, North Carolina) Distribution: USA (NC); new combination ex Zyras, Klimaszewski et al. 2005

cinctipennis Eppelsheim, 1884a: 12 ( Myrmedonia, TL : Talysch bei Rasano) Distribution: Azerbaijan, Tajikistan, Uzbekistan, Turkey

= esau Dvořák, 1984: 197 ( Zyras (Pella), TL : UdSSR, Tadjikistan, Javroz bei Dušanbe) synonymy in Maruyama, 2006:142

cognata Märkel, 1842: 142 ( Aleochara, TL : Saxon. Mont.; HA: HA: Lasius (Dendrolasius) fuliginosus ) Distribution: Europe, Russia (European), Georgia, Turkey comes Sharp, 1874: 10 ( Myrmedonia, TL : Japan; HA: Lasius (Dendrolasius) fuji , L. (D.) spathepus , L. (D.) orientalis , L. (D.) nipponensis , L. (D.) capitatus ) Distribution: Japan cooterorum Maruyama, 2006a: 77 ( Pella, TL : Xiaolongmen (1400m), Dongling Yan-shan Beijing China; HA: Lasius (Cautolasius) fuji ) Distribution: China (Beijing region, Yunnan)

coreana Maruyama, 2006a: 125 ( Pella, TL : Korea: Seoul, Nowon, Sanggye-dong, Mt. Suraksan) Distribution: Korea criddlei Casey, 1911: 72 ( Myrmedonia, TL : Aweme, Manitoba) Distribution: Canada (Manitoba); new combination ex Zyras, Klimaszewski et al. 2005

discolor Assing, 2008a: 236 ( Pella, TL : TR, Kahramanmaras, Gezbeli Gec, 2000m, N38º12'14, E35º59'39; HA:

Camponotus sp. ) Distribution: Turkey ecuamazzonica Pace, 2008b: 347 ( Zyras (Pella), TL : Ecuador, Napo R., Sacha Lodge, 250m) Distribution:

Ecuador erratica Hagens, 1863: 118 ( Myrmedonia, TL : Germany; HA: Tapinoma erraticum ) Distribution: Austria,

Belgium, Bosnia & Hercegovina, Czech Republic, France, Germany, Italy (Sicilia), Romania, Slovakia, Spain,

Algeria, Morocco, Tunisia, Turkey

= ehlersi Eppelsheim, 1884: 365 ( Myrmedonia, TL : Morocco) synonymy in Fenyes, 1920: 297

= mustela Rottenberg, 1870: 25 ( Myrmedonia, TL : bei Catania) synonymy in Fenyes, 1920: 297 excepta Mulsant & Rey, 1861: 366 ( Myrmedonia, TL : France) Distribution: France, Spain fauveli Sharp, 1883: 199 ( Myrmedonia, TL : North America, Unites States - Mexico, Jalapa) Distribution: USA

(California, Louisiana, Texas), Mexico funesta Gravenhorst, 1806: 54 ( Aleochara, TL : Berolin [= Berlin], Germany; HA: Lasius (Dendrolasius)

fuliginosus) Distribution: Europe, Russia (European), Georgia, Algeria

= atrata Heer, 1839: 309 ( Myrmedonia, TL : Bern) synonymy in Mulsant & Rey, 1873: 63

= crassicornis Stephens, 1832: 113 ( Aleochara, TL : Suffolk; preoccupied, not Aleochara crassicornis

Gyllenhal, 1827) synonymy in Fenyes, 1920: 297 gesneri Klimaszewski, 2005: 29 ( Pella, TL : Acadia Research Forest, Sunbury Co., New Brunswick, Canada)

Distribution: Canada gibbera Assing, 2007d: 40 ( Pella, TL : TR, Izmir, WSW Hamamköy, Murtat Daği, 1230 m) Distribution: Turkey glooscapi Klimaszewski & Majka, 2009: 37 ( Pella, TL : Canada, Nova Scotia, Upper Rawdon, Hants Co.)

Distribution: Canada hampei Kraatz, 1862: 267 ( Myrmedonia, TL : Croatia; HA: Liometopum microcephalum ) Distribution: Albania,

Austria, Bulgaria, Czech Republic, Greece, Hungary, Slovakia, Serbia, Turkey hlavaci Maruyama, 2006a: 102 ( Pella, TL : Xiaolongmen (1400m), Dongling, Yan-shan, Beijing, China; HA:

Lasius (Dendrolasius) fuji ) Distribution: China (Beijing) horii Maruyama, 2006a: 37 ( Pella, TL : Japan, Hebi-numa, Teshio-gawa, Teshio-chô, Hokkaidô; HA: Lasius

(Cautolasius) flavus ) Distribution: Japan humeralis Gravenhorst, 1802: 90 ( Aleochara, TL : Berolin [= Berlin], Germany; HA: Formica rufa , F. pratensis ,

Lasius (Dendrolasius) fuliginosus ) Distribution: Europe, Turkey, Russia (European, Siberia), Georgia,

Kazakhstan

= grandis Motschulsky, 1844: 814 ( Myrmedonia, TL : Daurica) iberica Maruyama, 2006a: 109 ( Pella, TL : E Andalusien (MA), Sierra de Palmitera, SO Ronda) Distribution:

Spain indiscreta Sharp, 1888: 291 ( Myrmedonia, TL : Seba & Hakodate; HA: Lasius (Dendrolasius) fuji , L. (D.)

spathepus, L. (D.) nipponensis , L. (D.) capitatus ) Distribution: Japan (Hokkaidô, Honshû, Shikoku) intermedia Maruyama, 2006a: 154 ( Pella ; TL: Russia, Primorskyi Krai, Nadezhdunskyi, Kravtzovka; HA: Lasius

(Dendrolasius) fuji) Distribution: far east Russia japonica Sharp, 1888: 290 ( Myrmedonia cognata var. japonica ; TL: Bukenji, Japan; HA: Lasius (Dendrolasius)

fuji, L. (D.) spathepus , L. (D.) nipponensis ) Distribution: Japan, Korea, far east Russia jelineki Maruyama, 2006a: 86 ( Pella, TL : Tunis, El Kef) Distribution: Tunisia jureceki Dvořák, 1981: 55 ( Zyras (Pella), TL : Siberia or., Ussuria, Vladivostok; HA: Lasius (Dendrolasius) fuji ,

L. (D.) nipponensis , L. (D.) orientalis ) Distribution: China (Beijing region, Korea, east Russia

= micropterus Pace, 1998: 976 ( Zyras (Pella), TL : China, Beijing, Xiaolongmen) synonymy in Maruyama,

2006a: 70 kidaorum Maruyama, 2006a: 73 ( Pella, TL : Taihei, Maruseppu-chô, Hokkaidô, Japan; HA: Lasius (Dendrolasius)

fuji, L. (D.) spathepus , L. (D.) orientalis , L. (D.) nipponensis , L. (D.) capitatus ) Distribution: Japan (Hokkaido) kinomurai Maruyama, 2006a: 96 ( Pella, TL : Isshiki, Shoukawa-mura [=Shôkawa-mura] Gifu Pref., Japan; HA:

Lasius (Dendrolasius) fuji , L. (D.) spathepus , L. (D.) capitatus ) Distribution: Japan (Hokkaido, Honshû) kirgisica Assing, 2009a: 204 ( Pella, TL : Kyrgyzstan, Tschui, U. Bischkek, Boz Pel’dek, N 42 o 45’47’’, E

74 o 33’51’’) Distribution: China (Hunan) kishimotoi Maruyama, 2006a: 117 ( Pella, TL : Dahong-cun (1030m), Da’an Xiang, Longshan, Hunan Sheng,

China; HA: L. (D.) spathepus ) Distribution: China (Hunan) kuluensis Cameron, 1939: 545 ( Zyras (Pella), TL : Kulu: Naggar, 5000 feet) Distribution: India (Himachal

Pradesh) laeviceps Eppelsheim, 1880: 459 ( Myrmedonia, TL : Suramgebirge & Alexanderhilf; HA: Lasius (Lasius) cf.

emarginatus ) Distribution: Armenia, Azerbaijan, Georgia, Russia (south European region), Turkey laocaiensis Pace, 2008e: 11 ( Zyras (Pella), TL : Vietnam, Lao Cai, ca. 40 km SE Sapa, 1400m) Distribution:

Vietnam. laticollis Märkel, 1844: 203 ( Aleochara, TL : Saxon. Mont.; HA: Lasius (Dendrolasius) fuliginosus ) Distribution:

Europe, Turkey leonhardi Bernhauer, 1912e: 110 ( Zyras, TL : Sicilien, Monte Cola) Distribution: Italy (Sicily) limbata Paykull, 1789: 54 ( Staphylinus, TL : Sweden; HA: Lasius (Lasius) brunneus , L. (Cautolasius) flavus , L.

(Dendrolasius) fuliginosus ) Distribution: Europe, Russia (Siberia), Kazakhstan

= laevis Gravenhorst, 1802: 95 ( Aleochara, TL : [Brunswick, Germany]) synonymy in Mulsant & Rey, 1873: 55

loricata Casey, 1894: 324 ( Myrmedonia, TL : Grimsby, Canada & Ohio) Distribution: USA (OH), Canada

(NS,ON)

= cremastrogastris Wasmann, 1894a: 207 ( Myrmedonia, TL : Ross County, Ohio) synonymy in Klimaszewski et al. 2005: 721 lugens Gravenhorst, 1802: 95 ( Aleochara, TL : Berolin [= Berlin]; HA: Lasius (Dendrolasius) fuliginosus , L.

(Lasius) brunneus ) Distribution: Europe, Kazakhstan, Turkmenistan, Uzbekistan, Armenia, Turkey

= kawalli Hochhuth, 1872: 97 ( Myrmedonia, TL : Volh. [Volhynien] & K. [Kiew]), synonymy in Gusarov,

1991: 4 masakoae Maruyama, 2006a: 156 ( Pella, TL : Takao-san, Hachiôji-shi, Tokyo; HA: Lasius (Dendrolasius)

spathepus, L. (D.) fuliginosus , L. (Chthonolasius) umbratus ) Distribution: Japan (Hokkaido, Honshû, Shikoku),

Korea maura Fauvel, 1898: 110 ( Myrmedonia, TL : Provinces d‘Alger, Milianah, provinces de Constantine, Edough)

Distribution: Algeria, Morocco, Italy (Sicily)

= ragusae Fairmaire, 1898: 110 ( Myrmedonia, TL : Madonna dell'Alto sulla Madonie, 1700m) synonymy in

Bernhauer & Scheerpeltz, 1926: 701

plutenkoi Maruyama, 2006a: 127 ( Pella, TL : Russia, Primorie/S Artyom town env., 100 – 300 m, Ozemyi

Kluyich) Distribution: far east Russia primorskyana Maruyama, 2006a: 100 ( Pella, TL : Russia, Primorskyi Krai, Khasanskyi, Pluostrov Gamov, Bukhta

Vityaz; HA: Lasius (Dendrolasius) fuji ) Distribution: east Russia puetzi Assing, 2009a: 206 ( Pella, TL : China: Yunnan, Nujiang Lisu Aut. Pref., Gaoligong Shan, side valley 19 km

NW Liuku, 25 o 59’02’’ N, 98 o 42’23’’E) Distribution: China (Yunnan) pumila Fiori, 1914: 105 ( Zyras, TL : al pied di un Cerro nel-Bosco Umbria, Gargano; HA: Liometopum microcephalum ) Distribution: Italy rambouseki Bernhauer, 1929e: 192 ( Zyras, TL : Ostsibirien; HA: Lasius (Dendrolasius) fuji , L. (D.) spathepus , L.

(D.) orientalis, L. (D.) nipponensis ) Distribution: east Russia, Korea recisa Casey, 1911: 74 ( Myrmedonia, TL : Pennsylvania) Distribution: USA (PA) reuteri Assing, 2008a: 241 ( Pella, TL : N-Irak, N Mosul - SW Hakkari, Al-Amadiya, 1200m) Distribution: Iraq ruficollis Grimm, 1845: 126 ( Myrmedonia, TL : Steyermark; HA: Liometopum microcephalum ) Distribution:

Austria, Bulgaria, Croatia, France, Hungary, Italia, Macedonia, Romania, Slovakia, Turkey

= fernandi Fairmaire, 1855: 309 ( Myrmedonia, TL : Naples) synonymy in Mulsant & Rey, 1873b: 73

schillhammeri Maruyama, 2006a: 93 ( Pella, TL : Seitengraben des Piwnaja-Rjetschka Tales b. Wladivostok; HA:

Lasius (Dendrolasius) sp.) Distribution: east Russia schmitti Hamilton, 1895: 346 ( Myrmedonia, TL : St. Vincent [Pennsylvania]; HA: Formica subsericea )

Distribution: Canada (Ontario, Quebec), USA (Massachusetts, Pennsylvania), new combination in

Klimaszewski et al., 2005: 722 similis Märkel, 1845: 200 ( Aleochara, TL : Gegend von Meissen [& other unnamed localities, in Germany?]; HA:

Lasius (Dendrolasius) fuliginosus , Liometopum microcephalum ) Distribution: Europe, Turkey socia Sharp, 1874: 10 ( Myrmedonia, TL : Japan; HA: Formica japonica ) Distribution: Japan spreta Sharp, 1888: 291 ( Myrmedonia, TL : Sapporo and Hakodate; HA: Lasius (Dendrolasius) fuji , L. (D.)

spathepus, L. (D.) capitatus ) Distribution: Japan (Hokkaido, Honshû) sultanica Assing, 2008a: 238 ( Pella, TL : Umg. Akschehir, Petrowitz-Ressl Reise nach Kleinasien 1960)

Distribution: Turkey zhoui Maruyama, 2006a: 164 ( Pella, TL : Xiaolongmen (1400m), Dongling, Yan-shan, Beijing, China; HA: Lasius

(Dendrolasius) fuji, L. (D.) nipponensis ) Distribution: China (Beijing region), Russia (Far East),