Hastulopsis excostellata ( Sacco, 1891a )

Harzhauser, Mathias & Landau, Bernard M., 2023, The auger snails (Gastropoda, Conoidea, Terebridae) of the Miocene Paratethys Sea, Zootaxa 5385 (1), pp. 1-70 : 19-21

publication ID	https://doi.org/ 10.11646/zootaxa.5385.1.1
publication LSID	lsid:zoobank.org:pub:7FBEC9FB-E31E-4CA4-8BD0-BE6D35322C3E
DOI	https://doi.org/10.5281/zenodo.10459900
persistent identifier	https://treatment.plazi.org/id/7C37CE5F-FFB2-AB21-FF2C-F941317EF9C3
treatment provided by	Plazi
scientific name	Hastulopsis excostellata ( Sacco, 1891a )
status

Treatment

Hastulopsis excostellata ( Sacco, 1891a)

Figs 3F View FIGURE 3 , 9A–F View FIGURE 9

Terebra striata Bast. View in CoL — Hauer 1837: 417 [non Hastula striata (de Basterot, 1825) ].

Terebra striata Bast. View in CoL — Hörnes 1848: 17 [non Hastula striata (de Basterot, 1825) ].

Terebra costellata Sow. View in CoL — Hörnes 1852: 134, pl. 11, fig. 30 [non Zeacuminia costellata (G.B. Sowerby I, 1846) View in CoL ].

Terebra Hörnesi Beyr. View in CoL — Beyrich 1854: 115 (partim).

Terebra (Acus) costellata Sow. View in CoL — Hoernes & Auinger 1880: 108 [non Zeacuminia costellata (G.B. Sowerby I, 1846) View in CoL ].

* [ Terebrum (Terebrum)? Hoernesi] var. excostellata Sacc. — Sacco 1891a: 27 [pro Terebra costellata sensu Hörnes 1852 View in CoL : pl. 11, fig. 30].

? Hastula striata Basterot — Csepreghy-Meznerics 1954: 56 [non Hastula striata (de Basterot, 1825) ].

H [astula]. (H [astula].) striata striata (Bast.) — Sieber 1958: 162 [non Hastula striata (de Basterot, 1825) ].

T [erebra]. (T [erebra].) hoernesi Beyr. — Sieber 1958: 162 [non Strioterebrum hoernesi ( Beyrich,1854) ].

? Hastula striata (Basterot) 1825 — Bohn-Havas 1973: 1124 [non Hastula striata (de Basterot, 1825) ].

Type material. Holotype, NHMW 1851 View Materials /0006/0005, SL: 25.3 mm, MD: 6.3 mm, Baden ( Austria), illustrated in Hörnes (1852: pl. 11, fig. 30), Figs 9D View FIGURE 9 1 –D View FIGURE 1 2 View FIGURE 2 .

Illustrated material. NHMW 1851 View Materials /0006/0005a, SL: 29.5 mm, MD: 7.3 mm, Baden ( Austria), Figs 9A View FIGURE 9 1 –A View FIGURE 1 2 View FIGURE 2 . NHMW 1851 View Materials /0006/0005b, SL: 30.2 mm, MD: 7.2 mm, Baden ( Austria), Figs 9B View FIGURE 9 1 –B View FIGURE 1 2 View FIGURE 2 . NHMW 1851 View Materials /0006/0005c, SL: 28.8 mm, MD: 7.0 mm, Baden ( Austria), Figs 9C View FIGURE 9 1 –C View FIGURE 1 2 View FIGURE 2 . NHMW 2023 View Materials /0295/0003, SL: 29.2 mm, MD: 7.1 mm, Bad Vöslau ( Austria), Figs 9E View FIGURE 9 1 –E View FIGURE 1 2 View FIGURE 2 . NHMW 2023 View Materials /0295/0004, SL: 25.3 mm, MD: 5.9 mm, Bad Vöslau ( Austria), Figs 9F View FIGURE 9 1 –F View FIGURE 1 2 View FIGURE 2 . NHMW 2023 View Materials /0295/0005, SL: 12.5 mm, MD: 3.5 mm, Bad Vöslau ( Austria), Figs 3F View FIGURE 3 .

Additional material. 22 spec., NHMW 1869 View Materials /0001/0025, Bad Vöslau ( Austria) ; 101 spec., NHMW 2010 View Materials /0004/1089, Bad Vöslau ( Austria) ; 2 spec., NHMW 1866 View Materials /0001/0571, Bad Vöslau ( Austria) ; 11 spec., NHMW 1869 View Materials /0001/0066, Baden-Sooss ( Austria) ; 10 spec., NHMW 2010 View Materials /0004/1092, Bad Vöslau ( Austria) ; 5 spec., NHMW 1997 View Materials z10178/1203, Bad Vöslau ( Austria) ; 21 spec., NHMW 2013 View Materials /0078/0415, Baden ( Austria) ; 15 spec., NHMW 2010 View Materials /0004/1091, Bad Vöslau ( Austria) .

Revised description. Medium sized, slender conical shell of up to 11 teleoconch whorls; apical angle 15–18°. Protoconch broad conical of 2.75 convex whorls. Early teleoconch whorls flat sided with narrowly impressed suture. Sculpture of low, weakly opisthocyrt axial ribs, separated by interspaces of about equal width. Subsutural band narrow and subobsolete. Later whorls with faint concavity below faint narrow subsutural band. Axial ribs shallowly opisthocyrt, of variable density. Last whorl moderately high, ~35% of total height, slightly convex, moderately contracted at base. Fasciole prominent, convex with few stronger growth increments, adapically delimited by weak carina. Aperture moderately narrow pyriform. Columella twisted, strongly excavated in adapical half. No columellar fold. Anal canal weakly incised. Outer lip thin. Siphonal canal moderately long, narrow, twisted, deflected to the left, moderately notched at tip.

Discussion. The Ottnangian (Early Miocene) Hastulopsis fuchsii ( Hoernes, 1875) is very similar to H. excostellata but differs in its lower spire and a relatively higher last whorl. Hastulopsis pedemontana ( Sacco, 1891a) ( Sacco 1891a: pl. 1 fig. 61) and H. striatellata ( Sacco, 1891a) ( Sacco 1891a: pl. 1, fig. 62) are both based on single fragmentary specimens from the Miocene of the Colli Torinesi ( Italy) without more detailed locality information. Both species, which might be conspecific, are morphologically very close to H. excostellata and differ only in the denser axial sculpture. Due to the fragmentary preservation, it remains unclear if these species are conspecific with H. excostellata . Fully grown specimens of H. hoernesi ( Beyrich, 1854) , from the Middle Miocene of the North Sea, differ in their lower spire, weaker axial sculpture, and less convex last whorl (see Beyrich 1854: pl. 6, figs 13a–b, 14a–b; Janse & Janssen 1983: text fig. 10, pl. 3, fig. 19; Wienrich 2006: figs 3/4, 4/6–10). Hastulopsis exilis ( Bell, 1871) , from the Early Pliocene of the Loire Basin ( France), might be another closely related species, which differs in its smaller size, wider spaced axial ribs, the more convex whorls and the slight basal angulation of the last whorl (see Ceulemans et al. 2018: pl. 6, figs 5–9). Hastula striata (de Basterot, 1825) from the Burdigalian of France, is slenderer, has a higher last whorl and it forms axially elongated beads at the adapical and abapical suture (see Peyrot 1931: pl. 10, figs 49–50).

Paleoenvironment. The occurrence in the Baden Formation suggests middle to outer neritic water depths ( Kranner et al. 2021).

Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Baden, Bad Vöslau ( Austria) ( Hoernes & Auinger 1880);? Cserhát Mountains ( Hungary) ( Csepreghy-Meznerics 1954); Făget Basin: CoŞteiu des Sus ( Romania) ( Hoernes & Auinger 1880).

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Order	Neogastropoda
Family	Terebridae
Genus	Hastulopsis