Toxognathus coomani Fleutiaux, 1940

Rosa, Simone Policena, 2014, Phylogenetic analysis and taxonomic revision of Physodactylinae (Coleoptera, Elateridae), Papéis Avulsos de Zoologia (São Paulo) 54 (18), pp. 217-292 : 241-242

Toxognathus coomani Fleutiaux, 1940

( Figs. 8 , 34H )

Toxognathus coomani Fleutiaux, 1940a: 35 .

Redescription (male, 39H): Integument brown to dark brown, legs and antennae lighter. Total length 5.5-6.5 mm; elytral base 1.03-1.08x as wide as pronotum, elytra 2.65-2.67x longer than pronotum. Frons concave along the anterior margin; frontal carina complete. Antennae surpassing the posterior angle of pronotum by one antennomere; antennomeres IV-X as long as wide and convex. Mandible bidentate ( Fig. 8A ). Pronotum ( Fig. 8B ) flattened 1.11-1.16x wider than long with sides slightly rounded, convergent on anterior 1/4; lateral carina entirely lateroventral (not visible from dorsal view) ( Figs. 8C, D ), posterior angles subparallel and convex; posterior margin with a pair of lateral incisions adjacent to the posterior angles; punctures 0.5 diameter apart on disc, coalescent on lateral border. Elytral interstices convex. Free margin of metacoxal plate short, triangular; tibiae slightly widened apicad with dorsal apical angle truncate. Tergite IX with anterior margin bisinuous.

Aedeagus ( Figs. 8G, H ): Phallobase 0.37x total length of aedeagus, 1.2x longer than wide, length ratio between lateroposterior and median parts 3; paramere with ventral anteromedian margin acute; basal struts 0.28x total length of penis; ventral sclerite linear.

Holotype: [Ton Kin, (Lac Thô), Hoa Binh, A. de Cooman], [ Toxognathus coomani , type, Collection Fleutiaux], [Type], [Muséum Paris, Coll. E. Fleutiaux], [ Toxognathus coomani (Fleutiaux) , J. Chassain det. 05], [Lectotype], (MNHN).

Material examined: VETNAM. Tonkin, 4 exs. ( MNHN) .

Distribution: VIETNAM. Hoa Binh: Lac Thuy.

Remarks: Toxognathus coomani shares with T. mouhoti and T. fairmarei the bidentate mandibles. It is most similar to T. mouhoti by its body size, eye size, shape of prothorax, free margin of metacoxal plate short and triangular and widened tibiae. It can be distinguished mainly by its frons, which is concave along the anterior border, complete frontal carina, shorter antenna and a pair of short notches on posterior margin of pronotum.

Figures

FIGURE 8: Toxognathus coomani. A, mandible (ventral); B, C, D, prothorax (dorsal, ventral, lateral); E, hind wing; F, abdomen (ventral); G, H, aedeagus (dorsal, ventral).

FIGURE 34: Habitus. A, Margogastrius schneideri (13.0 mm); B, Teslasena femoralis (9.0 mm); C, T. foucarti (9.5 mm); D, Idiotropia henoni (4.5 mm); E, Oligostethius capensis (10 mm); F,Toxognathus bakeri (6.0 mm); G, T. beauchenei (9.0 mm); H, T. coomani (6.5 mm); I, T. costulatus (11.0 mm); J, T. doherty (8.0 mm); K, T. fairmairei (7.5 mm); L, T. mouhoti (5.0 mm); M, Dactylophysus hirtus sp. nov. (10.5 mm); N, D. capixabensis nom. nov. (9.0 mm); O, D. tibialis (12.0 mm); P,Heterocrepidius mendax (9.0 mm); Q, Physodactylus asper sp. nov. (9.5 mm); R, P. besckei (11.0 mm). A, D = lectotypes; E-G, I, J, L, M, P, Q = holotypes.