Anterhynchium (Anterhynchium) cariosum Giordani Soika, 1987

Selis, Marco & Carpenter, James M., 2023, Revision of the Afrotropical species of the genus Anterhynchium de Saussure (Hymenoptera: Vespidae: Eumeninae), Zootaxa 5233 (1), pp. 1-102 : 76-78

Anterhynchium (Anterhynchium) cariosum Giordani Soika, 1987 stat. nov.

( Fig. 23 )

Anterhynchium synagroides cariosum Giordani Soika 1987: 199 , ♁, ♀ —“ Somalia: Belet-Uen ” (holotype NMBZ).

Diagnosis. This species can be recognized by combination of following characters: coloration entirely black with red-orange antennae, posterior lobe of tegula not equaling parategula, dorsal faces of propodeum entirely punctured, lateral and inferior carinae of propodeum denticulate, posterior face of propodeum strongly striate, T2–T5 with deep and coarse punctures and interspaces reduced to spines and carinae, S2 without median longitudinal furrow and not depressed in the middle, clypeus of male deeply emarginate at apex and with distinct large punctures on whole surface, tyloids present only on apical half of antenna, male F11 claw-shaped, mid femur of male simple, vertex of female with foveae placed in barely recognizable depression.

Description. MALE. Fore wing length 10.5–15.0 mm.

Head 1.2 × as wide as long in frontal view. Clypeus in lateral view weakly convex; in frontal view 1.1 × as long as wide, apical margin with deep semicircular incision, incision 0.7× as deep as wide, apical teeth long and triangular, sharply pointed apically. Interantennal space carinate. Distance from posterior ocellus to occipital carina 2.4 × as long as the distance from posterior ocellus to inner eye margin. Gena as wide as eye at bottom of ocular sinus; occipital carina complete, shortly lamellate. Scape 3.7 × as long as apically wide; F1 2.2 × as long as wide; F2–F7 longer than wide; F8–F10 subquadrate; F11 digitiform, apically rounded and weakly curved in lateral view, apex reaching base of F9; F6–F10 with tyloids. Mandibles tridentate, apical tooth long and weakly curved apically, mid tooth forming a weakly convex cutting margin, inner tooth triangular; space between mid and inner teeth deep and wide, rounded at bottom. Pronotal carina complete, elevated into a very short lamella on sides of dorsal face; pretegular carina fine, preceded by a shallow depression on ventral half. Mesoscutum 0.9 × as long as wide between tegulae, distinctly convex in lateral view. Scutellum convex, with a median longitudinal furrow deeper on posterior half. Metanotum angled in lateral view, with a short horizontal face and a longer vertical face, faces separated by a crenulate margin depressed on middle line. Tegula elongate, outer margin weakly convex, posterio lobe triangular and short, not reaching apex of parategula. Parategula large and straight, dorsally convex, with rounded apex. Mesepisternum weakly convex, epicnemial carina complete and strong, becoming lamellate ventrally and forming an elevated angle on ventral corners. Propodeum in lateral view oblique; sides of posterior face flat and angularly meeting on median line, therefore posterior face large and weakly depressed; dorsal face convex; lateral face flat with apical depression; dorsal carinae absent, therefore dorsal and posterior faces are not clearly separated; lateral carinae strong and denticulate, with larger teeth on posterolateral angles of propodeum. T 1 in dorsal view 0.65 × as long as wide, almost entirely parallel-sided, in lateral view with a strongly convex vertical face which continues in a longer flat horizontal face; T 2 in dorsal view wider than long, in lateral view weakly convex, with preapical shallow depression; S 2 in lateral view regularly convex from base to apex.

Head with deep flat-bottomed punctures, interspaces as long as or shorter than punctures diameter on frons, wider on vertex and gena; interantennal space and lower third of ocular sinus smooth. Clypeus densely microreticulate and dull, with small deep punctures, interspaces several times as long as punctures diameter. Pronotum and mesoscutum with very dense and deep punctures, interspaces forming a narrow reticulation, some punctures confluent with each other; ventral corners of pronotum micropunctate with some elongate shallow punctures; punctures on mesoscutum becoming denser posterolaterally. Scutellum and horizontal face of metanotum roughly punctured, interspaces irregular and reduced to sharp carinae and spines; vertical face of metanotum densely micropunctate with some shallow large punctures dorsally. Mesepisternum, except epicnemium and posterior margin, roughly punctured, interspaces on dorsal half narrow and carine-like, becoming flat ventrally. Metaepisternum densely microreticulate, almost dull, with some short striae on anterior margin. Dorsal faces of propodeum roughly punctured, large flat-bottomed punctures with narrow and sharp interspaces; posterior face with dense oblique irregular striae, some large punctures on dorsal margin; lateral face micropunctate and dull, with sparse transverse shallow striae and large oblique punctures. T1 dull, with sparse shallow punctures on dorsal face; lateral margins coarsely punctured; T2 dull, with shallow and sparse oblique punctures, becoming deeper on sides, apical fourth depressed and very roughly punctured, with interspaces sharp and irregular, reduced to short carinae and spines; T3–T5 punctured like apical part of T2; T6 with smaller and sparser punctures than previous terga, interspaces larger than punctures diameter; T7 densely micropunctate and dull. S1 smooth and shiny on basal petiole, enlarged apical part transversely rugose with shallow flat-bottomed punctures on sides; S2 microreticulate, with very dense micropunctures on lateral thirds and deep oblique punctures, interspaces larger than punctures diameter; S3–S6 sculptured like S2, but micropunctures deeper and becoming larger on apical terga, macropunctures becoming less numerous; S7 sculpted similar to T7, but micropunctures larger and deeper. Whole body covered by short and dense black pubescence, some longer black setae on frons, mesepisternum, posterolateral angles of propodeum, S1 and base of S2.

Black; antennae ferruginous, weakly darkened above; last tarsomere of all legs ferruginous; wings strongly fuscous, with blue-purplish reflections.

FEMALE. Fore wing length 14.0–17.0 mm.

Like male, except: head 1.25 × as wide as long in frontal view; clypeus in frontal view short pyriform, as long as wide, apical margin narrow and deeply incised, incision as deep as wide, apical teeth triangular and pointing out, apically rounded; gena 1.1 × as wide as eye at bottom of ocular sinus; cephalic foveae placed in barely recognizable depression, not clearly distinct from rest of vertex; scape 3.1 × as long as apically wide; F1 2.4 × as long as wide; F10 bullet-shaped; mandible quadridentate; T 1 in dorsal view 0.85 × as long as wide; clypeus more densely punctured.

Distribution. Kenya *, Somalia, Tanzania * ( Giordani Soika 1987) ( Fig. 20 ).

Remarks. This species was described as a subspecies of Anterhynchium (A.) synagroide due to the almost identical appearance, but deserves to be raised to specific rank due to the following characters [characters of synagroide in brackets]: apical punctation of T2–T 4 in female and T2–T 5 in male extremely strong and confluent, with interspaces reduced to sharp carinae or spines [apical punctation not so conformed], S2 without median depression and basally entirely convex from side to side [S2 with shallow median depression and basal convexity interrupted medially], entirely black with orange antennae [black and dark red with apical metasomal segments bright orange].

Figures

FIGURE 20. Distributional map of synagroide species-group.

FIGURE 23. Anterhynchium cariosum Giordani Soika, 1987. A, habitus of male. B, head of male in frontal view. C, head of female in frontal view. D, head of female in dorsal view. E, flagellum of male in lateral view. F, T2 in dorsal view.