Quercus novomexicana (A.DC.) Rydb., Bull.

Bartholomew, Bruce & Almeda, Frank, 2023, Nomenclator botanicus of Fagaceae in Latin America, Phytotaxa 580 (1), pp. 1921-1935 : 1921-1935

Quercus novomexicana (A.DC.) Rydb., Bull. New York Bot. Gard. 2(6): 208 (1901).

Quercus stellata Wagenh. var. utahensis A.DC, Prodr. 16(2.1): 22 (1864). Type :— U.S.A. (Holotype: MO?). According to A. de Candolle (1864, p. 22) the type was collected between Salt Lake and the Sierra Nevada. The specimen was from the “(h. Engelm.!)” so it should be at MO .

Quercus undulata Torr. var. gambelii (Nutt.) Engelm., Trans. Acad. Sci. St. Louis 3: 382 (1876).

Quercus utahensis (A.DC.) Rydberg, Bull. New York Bot. Gard. 2(6): 202 (1901).

Distribution: Mexico (Chihuahua, Coahuila, Nuevo León, Sonora, Veracruz), U.S.A. (Arizona, Colorado, Nevada,

New Mexico, Oklahoma, Texas, Utah, Wyoming).

Quercus germana Schltdl. & Cham., Linnaea 5(1): 78 (1830). Type :— MEXICO, Veracruz: C. J. W. Schiede 21 (Holotype at B destroyed. Lectotype (designated here): HAL0060343 !; isolectotypes: GOET (image)!, HBG (image)!, LE!, M!) .

Quercus germana var. echinata Trel., Mem. Natl. Acad. Sci. 20: 52 (1924). Type:— MEXICO, Veracruz: L. Hahn 344 (Lectotype (designated here): P00754016 !). Trelease (1924, p. 52) did not indicate place of deposit for Hahn 344, but most of the Hahn specimens he cites were at B and now destroyed. The only specimen of Hahn that we found was at P; it was annotated by Trelease in 1913 as an isotype.

Quercus substenocarpa Trel., Mem. Natl. Acad. Sci. 20: 52, pl. 41 (1924). Type:— MEXICO, San Luis Potosí: C. G. Pringle 3969 (Holotype: BR0000006915407 !; isotypes: A!, BM!, E!, F!, G!, GH!, K!, MEL (image)!, MO!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!).

Distribution: Mexico (Coahuila, Guanajuato, Hidalgo, Jalisco, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis

Potosí, Tamaulipas, Veracruz).

Quercus glabrescens Benth., Pl. Hartw. 56 (1840). Type:— MEXICO, Hidalgo: T. Hartweg 428 (Lectotype (designated here): K000512603 !; isolectotypes: A!, BM!, CGE!, E!, FI!, G!, K!, L!, LD (image)!, LE!, NY!, OXF!, P!, UCSB (fragment)!). There are two sheets of Hartweg 428 at K in the Bentham Herbarium, one of which is here lectotypified.

Quercus ambigua Bonpl., Pl.Aequinoct. 2: 51, pl. 93 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4162 (Holotype: P00129745 !; isotypes: B-W (as s.n.)!, UCSB (fragment from P holotype)!).

Quercus bonplandiana Sweet, Hort. Brit. 370 (1826). This is an invalid name as the holotype is the same as Q. ambigua Bonpl.

Quercus glabrescens var. integrifolia Liebm. ex A.DC., Prodr. 16(2.1): 35 (1864). Type :— MEXICO, Veracruz: F. M. Liebmann 3488 (Lectotype: C 10012484! designated by Trelease (1924, p. 217, pl. 34b); isolectotypes: C!, CAS!, UCSB (fragment)!). A . de Candolle cited “Liebm.! h. Copenh. f. 165–167.” These numbers are three Cupulif. Lbm. sheet numbers for Liebmann 3488. Trelease effectively lectotypified sheet 166 at C . Sheet 167 is at C as well as a fragment at UCSB, and sheet 165 is now at CAS.

Quercus radlkoferiana Trel., Mem. Natl. Acad. Sci. 20: 49, pl. 35a (1924). Type:— MEXICO, Puebla: F. M. Liebmann 3483 (Holotype at B destroyed. Lectotype (designated here): C 10012485!; isolectotypes: C!, CAS!, GH!, M!, UCSB (fragment from C lectotype)!, UCSB (fragment)!). Trelease (1924, p. 49) indicated that “ Liebmann, 156, 3483 ” is the type, so the specimen of Cupulif. Lbm. 156 at C would apparently be the holotype. However, in the caption of pl. 35a on p. 217 Trelease wrote “The type Mexico, Liebmann, 156 ” and indicates that this image was taken at B. It would seem more likely that the specimen at B would have had just the number 3483 on it, which is an assigned number for a particular collection, rather than just Liebmann 156 which is an assigned number for individual sheets. Because the specimen at B was destroyed, and the label is not shown in Trelease’s pl. 35a, it is impossible to know for certain what was on the B label. For this reason, we are lectotypifying the C specimen Cupulif. Lbm. 156.

Distribution: Mexico (Chiapas, Ciudad de México, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán,

Morelos, Oaxaca, Puebla, San Luis Potosí, Sinaloa, Tlaxcala, Veracruz, Zacatecas). The occurrence of Q. glabrescens in Querétaro and Tamaulipas reported by Villaseñor (2016) needs to be confirmed.

Quercus glaucescens Bonpl., Pl. Aequinoct. 2: 29, pl. 78 (1809). Type:— MEXICO, Guerrero: A. Bonpland 3921 (Holotype: P00129726 !; isotypes: B-W!, CGE (as s.n.)!, UCSB (fragment from P holotype)!). Although the specimen at P is marked as an isotype, it is actually the holotype and has a description in Bonpland’s hand.

Quercus chinantlensis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 179 (1854) . Type :— MEXICO, Oaxaca: F. M. Liebmann 3471 (Lectotype: C 10012469! designated by Trelease (1924, p. 41, pl. 28b); isolectotypes: C!, CAS!) .

Quercus cuneifolia Liebm. non Raf. (1838), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 182 (1854). See Q. pinalensis below.

Quercus nigrirhachis Trel., Repert. Spec. Nov. Regni Veg. 33: 314 (1934). Type:— MEXICO, Guerrero: L. Schultze 493 (Holotype: at B destroyed). The determination of Q. nigrirhachis as probably Q. glaucescens is based on the Trelease (1934) description as no images or original specimens exist.

Quercus pinalensis Trel., Mem. Natl. Acad. Sci. 20: 46, pl. 28a (1924). Type:— MEXICO, Sinaloa: B. Seemann 1456 (Holotype: K000512594 !; isotypes: BM!, K!). Nom. nov. for Q. cuneifolia Liebm. (1854) non Raf. (1838). The holotype is the K specimen with the determination by Liebmann in his hand.

Quercus synthetica Trel., Mem. Natl. Acad. Sci. 20: 151, pl. 297a (1924). Type:— MEXICO, Oaxaca: A. Ghiesbreght 121 (Holotype: C 10012534!; isotypes: P!, UCSB (fragment from C holotype)!, UCSB (fragment)!).

Quercus synthetica var. crenifolia Trel., Mem. Natl. Acad. Sci. 20: 151, pl. 296 (1924). Type:— MEXICO, Veracruz: A. Ghiesbreght 117 (Holotype: P00754020 !; isotypes: P!, UCSB (fragment)!).

Quercus texquitzinae Trel., Repert. Spec. Nov. Regni Veg. 33: 315 (1934). Type:— MEXICO, Guerrero: L. Schultze 32 (Holotype: at B destroyed). We suspect that Q. texquitzinae is probably Q. glaucescens based on the Trelease (1934) description as no images or original specimens exist.

Distribution: Mexico (Colima, Guerrero, Jalisco, México, Michoacán, Nayarit, Oaxaca, Puebla, Querétaro, San Luis

Potosí, Sinaloa, Tamaulipas, Veracruz).

Quercus glaucoides M.Martens & Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 209 (1843). Type :— MEXICO, Oaxaca: H. Galeotti 103 (Lectotype: BR0000009816220 !designated by Trelease (1924 p. 217.pl. 42);isolectotypes: BR!, G!, K!, P!, UCSB (fragment from BR lectotype)!, UCSB (fragment)!). M . Martens & Galeotti cite only one collection number, two sheets of which are at BR ; either one could be considered the type. Breedlove annotated the better of these two specimens as the holotype in 1987, but Trelease had already effectively lectotypified the other specimen. The specimen which is the lectotype would be a much better specimen had C. H . Muller not removed a small branch, apparently in 1958, which is now at UCSB .

Quercus baldoquinae Trel., Mem. Natl. Acad. Sci. 20: 53, pl. 43b (1924). Type:— MEXICO, Michoacán: R. Endlich 1354 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 43b)!). No extant original material exists, so we are lectotypifying the name on the image of the holotype.

Quercus cancellata Trel., Mem. Natl. Acad. Sci. 20: 55, pl. 49a (1924). Type:— MEXICO, Puebla: H. Schenk 236 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 49a)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus conjugens Trel., Mem. Natl.Acad. Sci. 20: 55, pl. 48 (1924). Type:— MEXICO, Guanajuato: C. G. Pringle 8841 (Lectotype (designated here): G00410068 !; isolectotypes: A!, BM!, CAS!, E!, F!, FI!, G!, GH!, HBG (image)!, ILL!, K!, L!, M!, MEXU!, MO!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!). Trelease (1924) indicated that the type was at B, now destroyed, and at G.

Quercus cordata M.Martens & Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 211 (1843). Type:— MEXICO, Oaxaca: H. Galeotti 111 (Holotype: BR0000009816237 !; isotype: UCSB (fragment from BR holotype)!).

Quercus glaucophylla Seemen, Bot. Jahrb. Syst. 29(1): 95 (1900). Type:— MEXICO, Oaxaca: C. G. Pringle 4843 (Holotype: BR0000009816374 !; isotypes: A!, BM!, CAS!, E!, ENCB (image)!, G!, GH!, K!, KFTA (image)!, MEL (image)!, MEXU!, MO!, NDG (image)!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!).

Quercus glaucophylla f. tlacolulana Trel., Mem. Natl. Acad. Sci. 20: 53 (1924). Type :— MEXICO, Oaxaca: C. Seler & E. G . Seler 1752 (Holotype at B destroyed. Lectotype (designated here): GH00062248 !) .

Quercus harmsiana Trel., Mem. Natl. Acad. Sci. 20: 54, pl. 45b (1924). Type:— MEXICO, Michoacán: R. Endlich 1356 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 45b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus mixtecana Trel., Mem. Natl. Acad. Sci. 20: 54 (1924). Type:— MEXICO, Oaxaca: C. Conzatti 1416 (Holotype at B destroyed. Lectotype (designated here): GH00969954 !; isolectotype: A!). No extant specimens of the type number, Schenck 235, from Puebla have been found, so the lectotype is based on the paratype, Conzatti 1416, from Oaxaca .

Distribution: Mexico (Chiapas, Chihuahua, Ciudad de México, Coahuila, Durango, Guanajuato, Guerrero, Hidalgo,

Jalisco, México, Michoacán, Morelos, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Tlaxcala, Veracruz ).

Quercus grahamii Benth., Pl. Hartw. 57 (1840). Type :— MEXICO: G. J. Graham 326 (Holotype: K000512981 !; isotypes: BM!, CGE!, GH!) .

Quercus acutifolia Neé f. abrupta Trel., Mem. Natl. Acad. Sci. 20: 192 (1924). Type:— MEXICO, Colima: H. Ross 534 (Lectotype (designated here): M!). Although Trelease (1924, p. 192) cited Ross 534, he did not specify the herbarium.

Quercus grahamii f. brevipes Trel., Mem. Natl. Acad. Sci. 20: 190, pl. 384a (1924). Type :— MEXICO, Veracruz: F. M. Liebmann 3437 (Holotype: C 10012490!; isotypes: CAS!, K (as s.n.)!, P (as s.n.)!). The caption of Trelease’s plate (1924, p. 229, pl. 384a) states “The type (Mexico, Liebmann, 33=3437). Photographed by permission at Kew.” The photograph in the plate of Liebmann 3437 (Cupulif. Lbm. 33) is of the specimen at C .

Quercus grahamii var. nelsonii Trel., Mem. Natl. Acad. Sci. 20: 190, pl. 384b (1924). Type:— MEXICO, Oaxaca: E. W. Nelson 533 (Holotype: US 00089534!; isotype: US!) .

Quercus xalapensis Bonpl. f. surculina Trel., Mem. Natl. Acad. Sci. 20: 193, pl. 393b (1924). Type :— MEXICO: W. F. Karwinski s. n. (Holotype: M!; isotype: UCSB (fragment from M holotype)!).

Distribution: Mexico (Colima, Guerrero, Hidalgo, Jalisco, México, Michoacán, Nuevo León, Oaxaca, Puebla,

Tamaulipas, Tlaxcala, Veracruz).

Quercus gravesii Sudw., Check List For. Trees U.S. 86 (1927). Type:— U.S.A., Texas: G. Sudworth s. n. (5 Nov 1913) (Holotype: A00034105!). Sudworth (1927) published Q. gravesii as a nom. nov. at the specific level for Q. texana var. chesoensis .

Quercus chesosensis (Sarg.) C.H.Mull., Amer. Midl. Naturalist 18(5): 850 (1937). This is an illegitimate name because Q. gravesii was already a nom. nov. at the specific level for Q. texana var. chesosensis .

Quercus coccinea Trel. var. microcarpa Torr., Rep. U.S. Mex. Bound., Bot. 2(1): 206 (1859). Type:— U.S.A., Texas: C. C. Parry et al. 1375 (Lectotype (designated here): NY00248625 !; isolectotypes: CAS!, UCSB (fragment)!, US!). There were two syntypes for this name.

Quercus cupreata Trel. & C.H.Mull., Bull. Torrey Bot. Club 63(3): 152 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 586 (Holotype: ILL00007521 !; isotypes: MEXU!, TEX (image)!, UCSB (fragment)!).

Quercus cupreata f. brachystachys C.H.Mull., J. Arnold Arbor. 17(3): 176 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 1327 (Holotype: A00055577 !; isotypes: MEXU!, NY!, P!, UCSB (fragment)!).

Quercus cupreata f. serrata Trel. & C.H.Mull., Bull. Torrey Bot. Club 63(3): 153 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 595 (Holotype: ILL00007519 !; isotypes: MEXU!, TEX (image)!, UCSB (fragment)!).

Quercus runcinatifolia Trel. & C.H.Mull., Bull. Torrey Bot. Club 63(3): 153 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 591 (Holotype: ILL00007571 !; isotype: UCSB (fragment)!).

Quercus runcinatifolia f. lata Trel. & C.H.Mull., Bull. Torrey Bot. Club 63(3): 153 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 593 (Holotype: ILL00007572 !; isotypes: TEX (image)!, UCSB (fragment)!).

Quercus shumardii Buckley var. microcarpa (Torr.) Shinners, Field & Lab. 24(1): 37 (1956).

Quercus stellapila (Sarg.) H.B. Parks ex Cory, Rhodora 38(455): 405 (1936).

Quercus tardifolia C.H.Mull., Bull. Torrey Bot. Club 63(3): 154 (1936). Type:— U.S.A., Texas: C. H. Muller s. n. (3 Jul 1932) (Holotype: ILL00007575 !; isotypes: CAS!, UCSB (fragment)!).

Quercus texana Buckley var. c hesosensis Sarg., Bot. Gaz. 65(5): 423 (1918). Type:— U.S.A., Texas: G. Sudworth s. n. (5 Nov 1913) (Holotype: A00034105 !). The protologue says 15 November but the date on the specimens is 5 November. This is also the type of Q. gravesii .

Quercus texana var. stellapila Sarg., Bot. Gaz. 65(5): 424 (1918). Type:— U.S.A., Texas: D. M. Andrews 74 (Lectotype (designated here): A00034106 !; isolectotype: A!). There are two specimens of Andrew 74 at A marked as the type by Sargent.

Distribution: Mexico (Chihuahua, Coahuila, Colima, Jalisco, Nuevo León, Tamaulipas), U.S.A. (Texas). The occurrence of Q. gravesii in Colima reported by Villaseñor (2016) needs to be confirmed.