Quercus greggii (A.DC.) Trel., Contr. U.S. Natl. Herb.

Bartholomew, Bruce & Almeda, Frank, 2023, Nomenclator botanicus of Fagaceae in Latin America, Phytotaxa 580 (1), pp. 1921-1935 : 1921-1935

Quercus greggii (A.DC.) Trel., Contr. U.S. Natl. Herb. 23(2): 185 (1922).

Quercus aculcingensis Trel., Contr. U.S. Natl. Herb. 23(2): 185 (1922). Nom. nov. for Q. reticulata Bonpl. var. crassifolia Oerst. (1869) non Q. crassifolia Benth. (1809) .

Quercus alpescens Trel., Mem. Natl. Acad. Sci. 20: 83, pl. 122b (1924). Type:— MEXICO: C. Ehrenberg 109 2 (Holotype at B destroyed. Lectotype (designated here): P00754039 !). Trelease (1924, p. 83) wrote “ Ehrenberg, 1091–3, Jan., 1840, the types” and on p. 219 for the caption of pl. 122b he wrote “The type (Mexico, Ehrenberg, 1093)” for a photograph of a specimen at B. The first statement indicates that there were three syntypes, but the second statement indicates that there is just the holotype. In either case all three specimens would have been at B and were destroyed.

Quercus loeseneri Trel., Mem. Natl. Acad. Sci. 20: 79, pl. 110b (1924). Type :— MEXICO, Coahuila: C. G. Pringle 10120 (Holotype at B destroyed. Lectotype (designated here): US 00089566!; isolectotypes: C!, E!, G!, GH!, HBG (image) !, ILL!, K!, L!, MEXU!, MO!, MP!, NY!, P !, PH!, UC!, UCSB (feagment) !, VT !).

Quercus reticulata Bonpl. var. crassifolia Oerst., Chênes Amér. Trop. 29, pl. 34 (10–16) (1869). Type:— MEXICO, Puebla: F. M. Liebmann 3548 (p.p. as to Cupulif. Lbm. 151) (Lectotype (designated here): C 10012458!). Trelease (1924, p. 219, pl. 110a) wrote “The types of Q. reticulata crassifolia (Mexico, Liebmann, 151).” The plate shows mostly Liebmann 3548 (Cupulif. Lbm. 151), plus a small part of Liebmann 3548 (Cuplulif. Lbm. 155) mounted on the same sheet, which were given different barcode numbers at C and were collected at different dates although from the same locality. Because Trelease wrote types rather than type, even though he mentioned Cupulif. Lbm. 151, he did not effectively lectotypify specimen Cupulif. Lbm. 151 which we are now lectotypfying. Because Liebmann 3548 (Cupulif. Lbm. 155) was collected on a different date, it cannot be considered an isolectotype.

Quercus reticulata var. greggii A.DC., Prodr. 16(2.1): 34 (1864). Type:— MEXICO, Coahuila: J. Gregg 380 (Holotype: G!; isotypes: GH!, MO!, UCSB (fragment)!).

Quercus revoluta Trel. f. dysophyllopsis Trel., Mem. Natl.Acad. Sci. 20: 79, pl. 112 (1924). Type:— MEXICO, Hidalgo: J. Berlandier 411 (Holotype: G00087110 !; isotypes: BM!, FI!, G!, P!, UCSB (fragment from G holotype)!, UCSB (fragment)!).

Distribution: Mexico (Chihuahua, Coahuila, Durango, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit,

Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tamaulipas, Tlaxcala, Veracruz, Zacatecas). The occurrence of Q. greggii in Nayarit reported by Villaseñor (2016) needs to be confirmed.

Quercus grisea Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 171 (1854) . Type:— U.S.A., Texas: C. Wright 665 (Holotype: K000832239 !; isotypes: G!, GH!, K!, MO!, NY!, P!, UCSB (fragment)!, US!) .

Quercus deserticola Trel., Mem. Natl. Acad. Sci. 20: 79, pl. 113 (1924). Type:— MEXICO, Hidalgo: C. A. Purpus 1233 (Holotype at B destroyed. Lectotype (designated here): GH02003320 !; isolectotypes: GH!, MO!, NY!, UC!, VT!). Neither an isotype of Uhde 309, nor duplicates of any of the Uhde paratypes from B have been found. The lectotype is designated from among the other paratypes that are the same species ( Q. grisea ) as the holotype photograph in Trelease (1924) pl. 113.

Quercus jaralensis Trel. f. berlandieri Trel., Mem. Natl. Acad. Sci. 20: 86, pl. 134 (1924). Type:— MEXICO, San Luis Potosí: J. Berlandier 1285 (Holotype: GH00062249 !; isotype: UCSB (fragment probably from GH holotype)!).

Quercus santaclarensis C.H.Mull., Amer. Midl. Naturalist 19(3): 583 (1938). Type:— MEXICO, Chihuahua: F. Shreve 7951 (Holotype: BH; isotypes: MICH!, UCSB (fragment from BH holotype)!, US!). An isotype in the protologue is also cited as being at the Desert Laboratory, Tucson , Arizona .

Quercus undulata Torr. var. grisea (Liebm.) Engelm., Trans. Acad. Sci. St. Louis 3: 393 (1877).

Distribution: Mexico (Aguascalientes, Chiapas, Chihuahua, Ciudad de México, Coahuila, Colima, Durango,

Guanajuato, Guerrero, Hidalgo, Jalisco, México , Michoacán, Morelos, Nuevo León, Oaxaca, Puebla, Querétaro, San

Luis Potosí, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas), U.S.A. (Arizona, New Mexico, Texas) .

Quercus hinckleyi C.H.Mull., Cont. Texas Res. Found., Bot. Stud. 1: 40, pl. 11 (1951). Type:— U.S.A., Texas: C. H. Muller & L. C. Hickley 8901 (Holotype: BH; isotypes: A!, BRIT (image)!, F!, LL (image)!, MO!, NY!, TEX (image)!, UC!, UCSB (fragment)!, US!) .

Distribution: Mexico (Chihuahua), U.S.A. (Texas).

Quercus hintonii E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 91 (1939). Type:— MEXICO, México: G. B. Hinton 6359 (Holotype: K000575008 !; isotypes: A!, BM!, F!, G!, K!, NY!, UCSB (fragment)!).

Quercus ochroesthes E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 94 (1939). Type:— MEXICO, México: G. B. Hinton 6576 (Lectotype (designated here): K000575014 !; isolectotypes: BM!, G!, GBH (image)!, K!, UCSB (fragment)!). There are two specimens at K, both of which are annotated by E.F.Warburg as “type.”

Distribution: Mexico (Guerrero, México, Michoacán).

Quercus hintoniorum Nixon & C.H.Mull., Brittonia 45(2): 147, fig. 1 (1993). Type :— MEXICO, Coahuila: R. M. Straw & M . Forman 1361 (Holotype: US; isotype: CAS!) .

Distribution: Mexico (Coahuila).

Quercus hirtifolia L.M.Vázquez, S.Valencia & Nixon, Brittonia 56(2): 137 (2004). Type:— MEXICO, Puebla: M.L.Vázquez 3069 (Holotype: MEXU (image)!; isotypes:BH, FCME).The specimen from Coahuila of C. B. Cowan 3566 at CAS! may possibly be a hybrid between Q. jonesii and Q. hirtifolia although there are no reported collections of Q. hirtifolia from Coahuila.

Distribution: Mexico (Hidalgo, Puebla, Veracruz).

Quercus huicholensis R.A.McCauley, Phytotaxa 471(3): 249. (2020). Type:— MEXICO, Jalisco: J. D. Bacon & R. D. Spellenberg 1475 (Holotype: NMC (NMC-065317) (image)!; isotypes: CAS (not yet received), CIIDIR, COLO, DAV, IBUG).Although we have not seen any of the type specimens, based on the description, illustrations, and the paratypes that we have seen, Q. huicholensis is very close to Q. urbani . Additional study is needed to determine whether they are different species.

Distribution: Mexico (Durango, Jalisco).

Quercus humboldtii Bonpl., Pl. Aequinoct. 2: 155, pl. 130 (1809). Type :— COLOMBIA, Cundinamarca: A. Bonpland 2083 (Holotype: P00669716 !; isotype: P!) .

Erythrobalanus duqueana O.Schwarz, Notizbl. Bot. Gart. Berlin-Dahlem 13: 495 (1937). Type:— COLOMBIA, Caldas: J. M. Duque 567 (Holotype: at B destroyed). The only specimen cited by Schwarz (1937) was Duque 567, but no isotypes have been found. There is a collection of Duque 1136 apparently from the same area, Río Nima, and also with the same determination at both F! and US!. Erythrobalanus duqueana is certainly Q. humboldtii but we have chosen not to neotypify the name based on Duque 1136 because it is not a recognized species.

Erythrobalanus tolimensis (Bonpl.) O.Schwarz, Notibl. Bot. Gart. Berlin-Dahlem 13: 496. 1937.

Quercus almaguerensis Bonpl., Pl. Aequinoct. 2: 157, pl. 131 (1809). Type:— COLOMBIA, Cuaca: A. Bonpland s. n. (Lectotype (designated here): image in Trelease (1924, pl. 269)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype. There is no doubt that Q. almaguerensis is Q. humboldtii , but no extant Bonpland collection has been found from Almaguer which is in Cauca Department, Colombia.

Quercus boyacensis Cuatrec., Notas Fl. Colombia 6: 37 (1944). Type:— COLOMBIA, Boyacá: J. Cuatrecasas & H. Garcia G. 9816 (Holotype: COL000000750 (image)!; isotypes: F!, UCSB (fragment)!, US!) .

Quercus colombiana Cuatrec., Notas Fl. Colombia 6: 37 (1944). Type:— COLOMBIA, Boyacá: E. Pérez A. & J. Cuatrecasas 8100 (Holotype: COL000000751 (image)!; isotypes: F!, P!, UCSB (fragment)!, US!) .

Quercus humboldtii var. lehmanniana Hieron. ex Trel., Mem. Natl. Acad. Sci. 20: 142, pl. 273b (1924). Type :— COLOMBIA, Antioquia: F. C. Lehmann 7453 (Holotype at B destroyed. Lectotype (designated here): K000648583 !) .

Quercus lindenii A.DC., Prodr. 16(2.1): 32 (1864). Type :— COLOMBIA, Tunja: J. Linden 1325 (Holotype: G00410065 !; isotypes: BM!, CGE!, G!, K!, P!, UCSB (fragment from G holotype)!, USCS (fragment)!).

Quercus tolimensis Bonpl., Pl. Aequinoct. 2: 153, pl. 129 (1809). Type:— COLOMBIA, Quindío: A. Bonpland 4165 (Holotype: P00669733 !; isotypes: P (as s.n.)!, UCSB (as s.n.) (fragment)!).

Distribution: Colombia, Panama.

Quercus hypoxantha Trel., Mem. Natl. Acad. Sci. 20: 170, pl. 339b (1924). Type:— MEXICO, Coahuila: C. G. Pringle 10227 (Holotype: GH00444446!; isotypes: ASU (image)!, CAS!, ENCB (image)!, LL (image)!, NY!, PH!, UC!, UCSB (fragment)!, UNM (image)!, US!, VT!, WIS!). Quercus hypoxantha forms a species complex in northern Mexico that has been studied by Nixon and Muller (1993). Several segregate species have been recognized. The complex could be treated in a broad sense as mostly a single species, but within this complex, Q. coahuilensis and Q. miquihuanensis are here treated as distinct.

Quercus errans Trel. f. graciliramis . C.H.Mull., J. Arnold Arbor. 17(3): 169 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 1273 (Lectotype (designated here): A00069935 !; isolectotypes: MEXU!, P!, UCSB (fragment)!). Muller (1936) designated two syntypes.

Distribution: Mexico (Chihuahua, Coahuila, Guanajuato, Nuevo León, Querétaro, San Luis Potosí, Tamaulipas,

Zacatecas). The occurrence of Q. hypoxantha in Guanajuato reported by Villaseñor (2016) needs to be confirmed.

Quercus iltisii L.M.González, Brittonia 55(1): 54, figs. 1, 3, 4 (2003). Type:— MEXICO, Jalisco: L. M. González V. et al. 4657 (Holotype: IBUG0160259 (image)!; isotypes: BH, CHAPA, ENCB, F (image)!, IEB, MEXU, MICH (image)!, NMC, NY (image)!, UC, US (image)!, WIS (image)!, ZEA).

Distribution: Mexico (Colima, Jalisco).

Quercus insignis M.Martens & Galeotti, Bull.Acad. Roy.Sci.Bruxelles 10(1): 219 (1843). Type:— MEXICO, Veracruz: H. Galeotti 125 (Lectotype: BR0000006912260 ! designated by Trelease (1924, p. 216, pl. 14); isolectotypes: G!, P!). H. Galeotti 123, 124, and 125 at BR are all syntypes, but Trelease (1924, p. 216) effectively lectotypified the name in the caption to his pl. 14.

Quercus davidsoniae Standl., Publ. Field Mus. Nat. Hist., Bot. Ser. 22(1): 14 (1940). Type:— PANAMA, Chiriquí: M. E. Davidson 864 (Holotype: F!; isotypes: A!, MO!, US!) .

Quercus insignis subsp. oocarpa (Liebm.) A.E.Murray, Kalmia 13: 26 (1983).

Quercus insignis var. oocarpa (Liebm.) A.E.Murray, Kalmia 13: 26 (1983).

Quercus insignis var. strombocarpoides (Liebm.) Oerst., Chênes Amér. Trop. 29, pl. H, pl. 28 (1869).

Quercus oocarpa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 184 (1854) . Type:— GUATEMALA: J. R. R. Warszewicz50 (Holotype at B destroyed. Lectotype (designated here): G00386111!; isolectotype: UCSB (fragment from G lectotype)!). Warszewicz 50a at B was both the holotype of Q. oocarpa and a syntype of Q. warsewiczii .

Quercus schippii Standl., Publ. Carnegie Inst. Wash. 461: 53 (1935). Type:— BELIZE, El Cayo: W. A. Schipp 1248 (Holotype: F (accn. 733679)!; isotypes: BM!, CAS!, G!, GH!, K!, LL (image)!, MICH!, MO!, NY!, S (image)!, UCSB (fragment)!).

Quercus seibertii C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 19, pl. 6, 7 (1942). Type:— PANAMA, Chiriquí: R. J. Seibert 225 (Holotype: A00055576 !; isotypes: K!, MO!, NY!, UCSB (fragment)!).

Quercus strombocarpa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 176 (1854) . Type:— MEXICO, Veracruz: F. M. Liebmann 3563 (Lectotype (designated here): C 10012531!; isolectotypes: C!, CAS!, GH!, M!, UCSB (fragment from C lectotype)!). Trelease (1924, p. 216) in the caption for pl. 17 wrote “Type material (Mexico foliage, Liebmann, 174=3563: fruit, Liebmann, Nov., 1841).” The Liebmann 3563 Cupulif. Lbm. 174 sheet at C is sterile, but presumably the acorn and cup also shown on pl. 17 are stored separately. Because he called this sheet type material rather than the type, we are lectotypifying the name based on this collection.

Quercus tomentocaulis C.H.Mull., Madroño 10(5): 130, pl. 9 (1950). Type:— HONDURAS, Morazán: L. O. Williams & A. Molina 12756 (Holotype: BH; isotypes: EAP (image)!, F!, GH!, MEXU!, MO!, UC!, UCSB (fragment from BH holotype)!, US!) .

Quercus warscewiczii Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 187 (1854) . Type:— PANAMA, Veraguas: B. Seemann 1230 (Lectotype (designated here): K000575035 !; isolectotype: BM!). Trelease (1924, p.43) had effectively lectotypified the type of Q. warscewiczii as Warscewicz 50a at B (destroyed) but this was also his lectotype of Q. oocarpa in the same publication, so we are relectotypifying the name based on a different syntype.

Quercus yunckeri Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 358 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6180 (Holotype: ILL00007538 !; isotypes: F!, G!, GH!, K!, MO!, NY!, S (image)!, UCSB (fragment)!).

Distribution: Belize, Costa Rica, Guatemala, Honduras, Mexico (Chiapas, Colima, Guanajuato, Guerrero, Hidalgo,

Jalisco, México, Michoacán, Nayarit, Oaxaca, Puebla, Tabasco, Veracruz), Nicaragua, Panama. The occurrence of Q .

insignis in Guanajuato, Hidalgo, México, and Michoacán reported by Villaseñor (2016) needs to be confirmed .

Quercus intricata Trel., Contr. U.S. Natl. Herb. 23(2): 185 (1922). Nom. nov. for Q. microphylla Née var. crispata A. DC (1864) non Q. crispata Steven. (1857) .

Quercus deliquescens C.H.Mull., Phytologia 42: 289 (1979). Type:— Mexico, Chihuahua: M. C. Johnston et al. 11388 (Holotype: TEX00370409 (image)!; isotype: UCSB (fragment)!).

Quercus derrumbaderoensis C.H.Mull., Amer. Midl. Naturalist 27(2): 471 (1942). Nom. nov. for Q. reticulata f. pungens C.H.Mull. (1936) non Q. pungens Liebm. (1854) .

Quercus intricata f. erratica Trel., Mem. Natl. Acad. Sci. 20: 85, pl. 128b (1924). Type:— MEXICO, Coahuila: C. G. Pringle 2862 (Holotype: CAS474678 !; isotypes: A!, VT!). Trelease (1924) in the caption to plate 128b states that Pringle 2862 is the type of Q. intricata f. erratica and that the image was photographed at St. Louis. The specimen of Pringle 2862 now at CAS was received as part of an exchange of duplicate type specimens with Harvard, and the GH specimen now at CAS was assumed at the time to be a duplicate isotype. However, the specimen now at CAS is the specimen in Trelease’s plate 128b, so it is the holotype of this form.

Quercus intricata f. ovata Trel., Mem. Natl. Acad. Sci. 20: 85, pl. 128a (1924). Type:— MEXICO, Coahuila: E. Palmer 746 (Holotype: MO-144164!; isotypes: GH!, NY!, UC!, UCSB (fragment from MO holotype)!, US!) .

Quercus microphylla Neé var. crispata A.DC., Prodr. 16(2.1): 36 (1864). Type:— MEXICO, Coahuila: J. Gregg 296 (Holotype: G00358136 !; isotypes: CAS!, GH!, UCSB (fragment)!).

Quercus reticulata Bonpl. f. pungens C.H.Mull., J. Arnold Arbor. 17(3): 166 (1936). Type:— MEXICO, Nuevo León: C. H. Muller 2417 (Holotype: A00055634 !; isotypes: CAS!, MEXU!, MO!, UCSB (fragment)!).

Quercus revoluta Trel. f. acuta C.H.Mull., J. Arnold Arbor. 17(3): 166 (1936). Type:— MEXICO, Nuevo León: C. H. Muller 2169 (Holotype: A00055628 !; isotypes: CAS!, MEXU!, MO!, UCSB (fragment)!).

Quercus supranitida C.H.Mull., Amer. Midl. Naturalist 27(2): 479 (1942). Nom. nov. for Q. revoluta f. acuta C.H.Mull. (1936) non Q. acuta Thunb. (1784) .

Distribution: Mexico (Chihuahua, Coahuila, Durango, Nuevo León, Oaxaca, San Luis Potosí, Tamaulipas, Zacatecas),

U.S.A. (Texas). The occurrence of Q. intricata in Oaxaca reported by Villaseñor (2016) needs to be confirmed.

Quercus invaginata Trel., Mem. Natl.Acad. Sci. 20: 87, pl. 137, 138a (1924). Type :— MEXICO, Coahuila: C. A. Purpus 5029 (Holotype at B destroyed. Lectotype (designated here): MO-144167!; isolectotypes: BM!, GH!, NY!, UC!, UCSB (fragment from MO lectotype)!, US!) .

Quercus invaginata f. purpusiana Trel., Mem. Natl. Acad. Sci. 20: 87, pl. 138b (1924). Type :— MEXICO, Coahuila: C. A. Purpus 5030 (Holotype at B destroyed. Lectotype (designated here): MO-144168!; isolectotypes: BM!, E!, F!, GH!, NY!, UC!, UCSB (fragment from MO lectotype)!, US!) .

Distribution: Mexico (Coahuila, Nuevo León, San Luis Potosí).

Quercus jonesii Trel., Mem. Natl. Acad. Sci. 20: 136, pl. 257b (1924). Type :— MEXICO, Jalisco: M. E. Jones 446 (Holotype: US00089543!; isotypes: RSA (image)!).

Quercus coccolobifolia Trel., Mem. Natl.Acad. Sci. 20: 136, pl. 258 (1924).Type:— MEXICO, Guanajuato: A. Bonpland 4269 (Holotype at B destroyed. Lectotype (designated here): P00754118!; isolectotype UCSB (fragment from P lectotype)!). The type collection number is not the same as the syntypes for Q. reticulata which are Bonpland s. n. but from the same locality. Trelease (1924, p. 136) was aware of the distinction and that Bonpland had used the specific epithet “cocolobaefolia” as an ined. name for what became the syntypes Q. reticulata . Martínez (1966) was the first person to synonymize Q. coccolobifolia under Q. jonesii .

Quercus endlichiana Trel., Mem. Natl. Acad. Sci. 20: 141, pl. 267b (1924). Type:— MEXICO, Durango: R. Endlich 6 (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 267b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus endlichiana f. minor C.H.Mull., J. Arnold Arbor. 17(3): 169 (1936). Type :— MEXICO, Nuevo León: C. H. Muller & M. T . Muller 1157 (Holotype: A00069873 !; isotype: P!) .

Quercus endlichiana f. serrata C.H.Mull., J. Arnold Arbor. 17(3): 170 (1936). Type :— MEXICO, Nuevo León: C. H. Muller & M. T . Muller 1330 A (Holotype: A00069871 !) .

Distribution: Mexico (Aguascalientes, Chihuahua, Coahuila, Durango, Guanajuato, Jalisco, Michoacán, Nayarit,

Nuevo León, Oaxaca, San Luis Potosí, Sinaloa, Sonora, Zacatecas). The occurrence of Q. jonesii in Oaxaca reported by Villaseñor (2016) needs to be confirmed.

Quercus kelloggii Newb., Pacif. Railr. Rep. 6(3): chapter 2, 28, 89, fig. 6 (1858). Nom. nov. at the specific level for Q. tinctoria Michx. var. californica . Newberry (1859, p. 28–29) cited Q. tinctoria var. californica as a synonym and indicated that he considered it to be neither Q. tinctoria nor Q. coccinea Münchh. although similar to both. He would also have been aware that Torrey (1857, p 138) thought that Q. tinctoria var. californica could be a new species when he wrote “it is probably a distinct species.” For these reasons we consider Q. kelloggii to be a nom. nov. for Q. tinctoria var. californica based on the same type (Bartholomew & Almeda 2022).

Quercus californica (Torrey) J.G.Cooper, Smithson. Rep. 1858: 261 . (1859). Nom. illeg. because Cooper cites Q. kelloggii Newb. (1857) as a synonym which is an older name.

Quercus sonomensis Benth. ex A.DC., Prodr. 60(2.1): 62 (1864). Type:— U.S.A., California: T. Hartweg 1966 (139) (Holotype: G00719751 (image)!; isotypes: GH (image)!, K (image)!, NY (image)!, P (image)!).

Quercus tinctoria W. Bartram var. californica Torr., Pacif. Railr. Rep. 4(5): 138. (1857). Type:— U.S.A., California: J. M. Bigelow s. n. (1854) (Lectotype: NY01470684 (image)! designated by Bartholomew & Almeda (2022, p. 28); isolectotype: GH (image)!). Torrey noted that “The greater part of the botanical collections made by D. J. M. Bigelow, in the Pacific Railroad Survey, under the charge of Captain Whipple, were submitted to me for examination in accordance with the instructions of the War Department.” Torrey also had available to him other collections which were in the Torrey Herbarium now at NY, including examples of the same oak made during the Wilkes Expedition of 1838–42, Frémont’s Expedition to California in 1845–47, and the Emory expeditions with a collection by C.C. Parry in 1850. It is clear that he relied on the collection localities of all these specimens when he wrote “It occurs throughout the valley of the Sacramento, and as far south as San Diego.” All of these NY specimens are syntypes including the collection J. M Bigelow made in the Napa Valley of California in 1854 which we are designating as the lectotype .

Distribution: Mexico (Baja California), U.S.A. (California, Oregon).

Quercus laceyi Small, Bull. Torrey Bot. Club 28(6): 358 (1901). Type:— U.S.A., Texas: H. Lacey s. n. (Holotype: NY). Breedlove considered Q. laceyi to be a synonym of Q. glaucoides , but Nixon & Muller (1992) convincingly show that they are different species.

Quercus breviloba (Torr.) Sarg. f. laceyi (Small) Trel., Mem. Natl. Acad. Sci. 20: 102 (1924).

Quercus breviloba subsp. laceyi (Small) A.Camus, Chênes Texte 2: 680 (1939).

Quercus glaucophylla f. lobata C.H.Mull., J. Arnold Arbor. 17(3): 163 (1936). Type :— MEXICO, Nuevo León: C. H. Muller & M. T . Muller 1281 (Holotype: A00055590 !; isotype: MEXU!) .

Quercus glaucophylla f. longifolia C.H.Mull., J. Arnold Arbor. 17(3): 163 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 589 (p.p.) (Holotype: A00055591 !). The specimen of C. H. Muller & M. T. Muller 589 at ILL is Q. calophylla , so it is not an isotype.

Quercus glaucophylla var. macropetiolata C.H.Mull., Bull. Torrey Bot. Club 63(3): 151 (1936). Type :— MEXICO, Nuevo León: C. H . Muller & M . T . Muller 574 (Holotype: ILL!; isotype: MEXU!) .

Quercus glaucophylla var. subrotundifolia C.H.Mull., J. Arnold Arbor. 17(3): 163 (1936). Type :— MEXICO, Nuevo León: C. H. Muller & M. T . Muller 1282 (Holotype: A00283983 !) .

Distribution: Mexico (Coahuila, Nuevo León, Tamaulipas), U.S.A. (Texas).

Quercus laeta Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 179 (1854) . Type:— MEXICO, Hidalgo: T. Hartweg 419 (Lectotype (designated here): K000512627!; isolectotypes: BM!, CAS!, CGE!, E!, G!, K!, NY!, OXF!, P!, UCSB (fragment)!). Liebmann cited syntypes of T. Hartweg 419 at B and K. Trelease (1924, p. 218, pl. 86b) effectively lectotypified the name based on Hartweg 419 at B. Because the B specimen was destroyed, we are re-lectotypifying the name based on the other syntype which is from the Hooker Herbarium at K and has a determination by Liebmann in his hand. Further study is needed regarding the relationship of Q. laeta and Q. arizonica . Breedlove believed that these two species might best be treated in a broad sense as a single species.

Quercus bipedalis Trel., Mem. Natl. Acad. Sci. 20: 89, pl. 143 (1924). Type:— MEXICO, Nayarit: J. N. Rose s. n. (10 Aug 1897) (Holotype: US 00089417!) .

Quercus clivicola Trel. & C.H.Mull., Bull. Torrey Bot. Club 63(3): 149 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 584 (Holotype: ILL00007560 !; isotypes: A!, MEXU!, TEX (image)!, UCSB (fragment)!).

Quercus clivicola f. consanguinea C.H.Mull., Bull. Torrey Bot. Club 63(3): 150 (1936). Type :— MEXICO, Nuevo León: C. H. Muller & M. T . Muller 597 (Holotype: ILL00007529 !; isotype: MEXU!) .

Quercus clivicola f. crenifolia Trel. & C.H.Mull., Bull. Torrey Bot. Club 63(3): 150 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 585 (Holotype: ILL00007528 !; isotypes: MEXU!, TEX (image)!, UCSB (fragment)!).

Quercus clivicola f. dentata Trel. & C.H.Mull., Bull. Torrey Bot. Club 63(3): 150 (1936). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 583 (Holotype: ILL00007527 !; isotypes: A!, CAS!, TEX (image)!, UCSB (fragment)!).

Quercus clivicola f. elongata C.H.Mull., J. Arnold Arbor. 17(3): 162 (1936). Type :— MEXICO, Nuevo León: C. H. Muller & M. T . Muller 1323 (Holotype: A00969952 !; isotypes: MEXU!, NY!, P!) .

Quercus laeta f. heterophylla Trel., Mem. Natl. Acad. Sci. 20: 71, pl. 87 (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 1009 (Holotype: P00754119 !; isotypes: P!, UCSB (fragment)!). Trelease (1924, p. 71) stated that the type is “Bourgeau, without number” at P, but the photograph in pl. 87, which Trelease designated in the caption to pl. 87 as the type, is of Bourgeau 1009 and also was annotated by Trelease as Q. laeta in 1913.

Quercus obscura Trel. non Seemen (1897), Mem. Natl. Acad. Sci. 20: 71, pl. 88 (1924). Type:— MEXICO: B.Seemann 1971 (Syntypes: C, GH00034124!, K000512630!; isosyntypes: BM!, UCSB (fragment)!). Because Q. obscura Trel. is a later homonym, we are not lectotypifying the name, but see below where we lectotypify the name Q. transmontana var. obscura based on the K specimen of Seemann 1971.

Quercus obscura Trel. non Seemen (1897) var. perpusilla Trel., Mem. Natl. Acad. Sci. 20: 71, pl. 89a (1924). Type:— MEXICO: B. Seemann 1976 (Lectotype (designated here): K000512631 !; isolectotypes: BM!, E (as s.n.)!, GH!). Trelease (1924, p. 71) cited a single collection but indicated that it was at both GH and K. In his pl. 89b he showed an image of the K specimen, but he did not indicate that it was “the type” as was his usual practice.

Quercus pallescens Trel., Mem. Natl. Acad. Sci. 20: 89, pl. 141b (1924). Type:— MEXICO, Jalisco: J. N. Rose 2960 (Holotype: US00089603!; isotypes: NY!, UCSB (fragment)!).

Quercus pandurata Bonpl. var. laeta (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 197 (1884).

Quercus prinopsis Trel., Mem. Natl. Acad. Sci. 20: 61, pl. 58b (1924). Type:— MEXICO, San Luis Potosí: C. A. Purpus 4941 (Holotype: MO-144170!; isotypes: BM!, E!, F!, GH!, P!, UC!, UCSB (fragment from MO holotype)!, US!) .

Quercus rossii Trel. f. arsenei Trel., Mem. Natl. Acad. Sci. 20: 180 (1924). Type:— MEXICO, Michoacán: G. Arsène 5913 (Lectotype (designated here): ILL00007610 !; isolectotypes: BR!, MEXU!, MO!, P!, US!). Trelease (1924, p. 180) cited three Arsène collections without indicating which is the type.

Quercus transmontana Trel., Mem. Natl.Acad. Sci. 20: 71, pl. 89b (1924). Type :— MEXICO, Michoacán: C. G. Pringle 4116 (Holotype at B destroyed. Lectotype (designated here): US 00089673!; isolectotypes: A!, BM!, BR!, CAS!, E!, F!, G!, GH!, K!, KFTA (image) !, MEL (image) !, MEXU!, MO!, NY!, P!, PH!, UC!, UCSB (fragment) !, US!, VT !).

Quercus transmontana f. lata (Trel.) A.Camus, Chênes Texte 2: 654 (1939).

Quercus transmontana var. lata Trel., Mem. Natl. Acad. Sci. 20: 72 (1924). Type:— MEXICO, Michoacán: G. Arsène 2975 (Syntypes at M lost. Lectotype (designated here): US00089672!; isolectotype: MPU!). Trelease (1924, p. 72) cited two Arsène syntypes at M that were destroyed. We are lectotypifying the name based on an isosyntype.

Quercus transmontana var. obscura A.Camus, Chênes Atlas 2: 97 (1935). Type:— Mexico: B. Seemann 1971 (Lectotype (designated here): K000512630!; isolectotypes: BM!, C!, GH (as s.n.)!, UCSB (fragment)!). Quercus obscura Trel. (1924) non Seemen (1897) is a later homonym, but the specific epithet of Trelease was used by A.Camus, based on the same type of which Trelease cited syntypes at C, GH, and K, as the basis for Q. transmontana var. obscura .

Distribution: Mexico (Aguascalientes, Baja California Sur, Chihuahua, Ciudad de México, Coahuila, Colima,

Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México , Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca,

Puebla, Querétaro, San Luis Potosí, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz, Zacatecas).

Quercus lancifolia Schltdl.&Cham., Linnaea 5(1):78(1830). Type:— MEXICO, Veracruz: C. J. W. Schiede 15 (Holotype at B destroyed. Lectotype: P00754120 ! designated by Valencia-A. & Coombes (2020, p. 89); isolectotypes: BM (as s.n.)!, P!, UCSB (fragment from P holotype)!, UCSB (fragment from BM as s.n.)!). The holotype at B would have been number 20. The species of oaks are numbered in accordance with the Linnaean treatment, 17 through 24. Other oaks were apparently numbered but not in this consecutive numbering system.

Quercus aaata C.H.Mull., U.S.D.A. Bur. Pl. Industr. Misc. Publ. 477: 27, pl. 23–26. (1942). Type:— GUATEMALA, Guatemala: P. C. Standley 58466 (Holotype: F0060002 F!; isotypes: F!, NA, UCSB (fragment)!, US) .

Quercus boqueronae Trel., Mem. Natl. Acad. Sci. 20: 48, pl. 33b (1924). Type:— MEXICO, Chiapas: C. A. Purpus 6997 (Holotype: ILL00009820 !; isotypes: BM!, F!, GH!, HBG (image)!, ILL!, MO!, NY!, UC!, UCSB (fragment)!, US!, VT!).

Quercus cyclobalanoides Trel., Proc. Amer. Philos. Soc. 54: 11, pl. 3 (1915). Type:— MEXICO, Chiapas: C. A. Purpus 6998 (Lectotype: ILL00007515 ! and ILL00010193 !designated by Trelease (1924, p. 216, pl. 20–21); isolectotypes: BM!, GH!, F!, ILL!, MO!, NY!, UC!, UCSB (fragment)!, US!). The lectotype is mounted on two sheets, one with just leaves and the other with just acorns and cups. The sheet with leaves has the collector’s label.

Quercus excelsa Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 174 (1854) . Type:— MEXICO, Veracruz: F. M. Liebamnn 3477 (Lectotype: C 10012479! designated by Trelease (1924, p. 47); isolectotypes: C!, CAS!, GH!, K (as s.n.)!, M!, UCSB (fragment)!). Trelease (1924, p. 47) wrote “ Liebmann, 205, 3477, Jan., 1943, the type)” by which he effectively lectotypified the Cupulif. Lbm. 205 sheet at C of Liebmann 3477. On p. 216 in the caption to pl. 29 he again wrote “The type (Mexico, Liebmann, 3477)” although in pl. 29 he shows photos taken of the two syntypes at C, neither of which are the Cupulif. Lbm. 205 sheet, so the caption to pl. 29 cannot be considered effective lectotypification. The three s.n. collections at K are almost certainly syntypes of Q. excelsa as they have the correct date and locality.

Quercus insignis M.Martens & Galeotti var. pilarius (Trel.) A.E.Murray, Kalmia 13: 26 (1983).

Quercus insignis subsp. pilarius (Trel.) A.E.Murray, Kalmia 13: 26 (1983).

Quercus lancifolia f. pilosiuscula Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 192 (1884). Type:— MEXICO, Veracruz: C. J. W. Schiede s. n. (Sep 1829) (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 32b)!). No extant original material exists, so we are lectotypifying the name based on the image of the holotype.

Quercus lancifolia var. monocarpa Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 192 (1884). Type:— MEXICO, Veracruz: F. M. Liebmann 3503 (Holotype: C 10012494!; isotypes: C!, CAS!, GH!, K (as s.n.)!, P!, UCSB (fragment)!). The holotype of Q. lancifolia var. monocarpa is also the lectotype of Q. leiophylla .

Quercus leiophylla A.DC., Prodr. 16(2.1): 71 (1864). Type:— MEXICO, Veracruz: F. M. Liebmann 3503 (Lectotype: C10012494 ! designated by Valencia-A. & Coombes (2020, p. 89); isolectotypes: C!, CAS!, GH!, K (as s.n.)!, P!, UCSB (fragment)!). Collections cited by A. de Candolle (1864) correspond to Liebmann 3505, some of which were collected in Oct 1841 and some in Nov 1841, but all at the same locality of S. Bartholome. A. de Candolle cites specimens as “ Q. lancifolia Liebm. ! in h. Copenh. et DC., non Cham. et Schl.” Of the six specimens originally at C, one is now at CAS. All six specimens were determined in A. de Candolle’s hand as Q. leiophylla . We have chosen one of the C specimens collected in Oct 1841 as the lectotype which is also the holotype of Q. lancifolia var. monocarpa Wenz.

Quercus leiophylla f. subintegra Trel., Mem. Natl. Acad. Sci. 20: 47 (1924). Type:— MEXICO, Veracruz: H. Galeotti 129 (Lectotype (designated here): G00358129 !; isolectotypes: BR!, G!, L!, P!, UCSB (fragment)!). Although Trelease (1924) cited a single specimen, he did not designate the herbarium.

Quercus molinae C.H.Mull., Madroño 10(5): 132, pl. 10 (1950). Type :— HONDURAS, Morazán: L. O . Williams & A . Molina 12753 (Holotype: BH; isotypes: EAP (image)!, F!, GH!, MEXU!) .

Quercus ovandensis Matuda, Amer. Midl. Naturalist 43(1): 198 (1950). Type:— MEXICO, Chiapas: E. Matuda 17835 (Holotype: MEXU00013001 !; isotypes: F!, MEXU!, UCSB (as s.n.) (fragment)!).

Quercus pilarius Trel., Mem. Natl.Acad. Sci. 20: 44, pl. 19 (1924). Type:— MEXICO, Chiapas: E. Reeves 11 (Holotype: ILL00007605 ! and ILL00010205 !). The holotype is mounted on two sheets. One sheet has just leaves and the other has the acorns and cups. The original label from the collection is on the sheet with just acorns and cups.

Quercus pilgeriana Seemen, Bull. Herb. Boissier , sér. 2, 4(7): 655 (1904). Type:— COSTA RICA, Alajuela: A. Tonduz 2197 (Holotype: BR0000006916176 !; isotypes: BR!, UCSB (fragment)!, US!). The label on the BR specimen indicates that it was collected by H. Pittier, but the protologue give the collection is by A. Tonduz 2197.

Quercus reevesii Trel., Mem. Natl. Acad. Sci. 20: 45, pl. 22 (1924). Type:— GUATEMALA, Quetzaltenango: E. Reeves s. n. (Feb 1918) (Holotype: ILL00007619 ! and ILL00010207 !; isotype: UCSB (fragment from ILL holotype)!). The holotype is mounted on two sheets. One sheet with just leaves and the other with just acorns and cups. The sheet with leaves has the label from the collector.

Quercus yousei Trel., Publ. Field Mus. Nat. Hist., Bot. Ser. 17(4): 358 (1938). Type:— HONDURAS, Comayagua: T. G. Yuncker et al. 6297 (Holotype: ILL00007539 !; isotypes: F!, G!, GH!, K!, MO!, NY!, S (image)!, UC!, UCSB (fragment)!, US!) .

Distribution: Belize, Costa Rica, El Salvador, Guatemala, Honduras, Mexico (Chiapas, Colima, Guerrero, Hidalgo,

Jalisco, México, Nayarit, Oaxaca, Puebla, San Luis Potosí, Tamaulipas, Veracruz), Nicaragua, Panama. The occurrence of Q. lancifolia in Colima, Nayarit, and Tamaulipas reported by Villaseñor (2016) needs to be confirmed.

Quercus laurina Bonpl., Pl. Aequinoct. 2: 32, pl. 80 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4143 (Holotype: P00129720 !; isotypes: CGE (as s.n.)!, G!, P!, UCSB (fragment)!).

Quercus acatenangensis Trel., Mem.Natl.Acad.Sci. 20: 163, pl. 320 (1924). Type :— GUATEMALA: J. R. R. Warszewicz 48 (Syntypes at B destroyed. Lectotype (designated here): K000512943 !; isolectotypes:G!, P!, UCSB (fragment)!). Nom. nov. for Q. longifolia Liebm. (1854) non Raf. (1838).

Quercus ambivenulosa Trel., J. Wash.Acad. Sci. 23(1): 62 (1933). Type:— GUATEMALA, Chimaltenango: A. F. Skutch 14 (Holotype: US 00089407!; isotypes: ILL!, US!) .

Quercus barbinervis Benth., Pl.Hartw. 56 (1840). Type:— MEXICO, Hidalgo: T. Hartweg 427 (Holotype: K000512950 !; isotypes: A!, BM!, CAS!, CGE!, E!, FI!, G!, K!, LD (image)!, LE!, NY!, P!).

Quercus bourgaei Oerst. ex Trel., Mem. Natl. Acad. Sci. 20: 168, pl. 336a (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 1013 (Holotype: G00358130 !; isotypes: BM!, BR!, C!, G!, GH!, K!, LE!, MPU!, P!, UCSB (fragment)!, US!) .

Quercus bourgaei var. ilicifolia Trel., Mem. Natl. Acad. Sci. 20: 168, pl. 336b (1924). Type:— MEXICO, Ciudad de México: C. G. Pringle 10315 (Lectotype (designated here): MO-144175!; isolectotypes: BM!, CAS!, F!, GH!, K!, ILL!, MEXU!, NY!, OKLA (image)!, P!, PH!, UC!, US!, VT!). Trelease (1924, p. 227) in the caption for plate 336 states that the type is at G, but because it could not be found there, we are lectotypifying an isotype specimen at MO which has a determination in Trelease’s hand.

Quercus caeruleocarpa Trel., Mem. Natl. Acad. Sci. 20: 163, pl. 321a (1924). Type:— MEXICO, Ciudad de México: R. Endlich 1365a (Holotype at B destroyed. Lectotype (designated here): image in Trelease (1924, pl. 321a)!). No extant original material exists, so we are lectotypifying the name based on an image of the holotype.

Quercus castanea Neé var. tridens (Bonpl.) A.DC., Prodr. 16(2.1): 72 (1864).

Quercus chrysophylla Bonpl., Pl. Aequinoct. 2: 42, pl. 87 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4062 (Holotype: P00669720 !; isotypes: B-W!, CGE (as s.n.)!, G!, P!, UCSB (fragment)!).

Quercus lanceolata Bonpl., Pl. Aequinoct. 2: 34, pl. 81 (1809). Type:— MEXICO, Guanajuato: A. Bonpland s. n. (Holotype: P00129724 !; isotypes: B-W!, G!, P!, UCSB (fragment)!).

Quercus lanceolata var. undulatodentata A.DC., Prodr. 16(2.1): 70 (1864). Type:— MEXICO, Veracruz: F. M. Liebmann 3496 (Lectotype (designated here): C 10012513!; isolectotypes: C!, CAS!). A.de Candolle (1864) cites two Liebmann collections at C by locality, not collection number. His citation of “In monte Orizaba alt. 8000 p.” corresponds to Liebmann 3496, and his citation of “prov. Oaxaca da Pico de Pelado alt. 9000 p.” corresponds to Liebmann 3495. Collections of both numbers at C are annotated in A.de Candolle’s hand. The placement of this variety needs additional study, but we consider the type to fall within the variation found within Q. laurina as broadly interpreted.

Quercus laurina var. barbinervis (Benth.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 205 (1884).

Quercus laurina var. ocoteifolia (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 205 (1884).

Quercus longifolia Liebm. non Raf. (1838), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 185 (1854). See Q. acatenangensis above.

Quercus malinaltepecana Trel., Repert. Spec. Nov. Regni Veg. 33: 318 (1934). Type:— MEXICO, Guerrero: L. Schultze 387 (Holotype: at B destroyed). No original material exists, so the only information is Trelease’s (1934) description. Quercus malinaltepecana might be synonymous with Q. laurina , but Breedlove thought that it might be Q. planipocula .

Quercus nitens M.Martens & Galeotti (1843) non Raf. (1838) var. ocoteifolia (Liebm.) A.DC., Prodr. 16(2.1): 69 (1864).

Quercus nitidissima Trel., Mem. Natl. Acad. Sci. 20: 122, pl. 218b (1924). Type:— MEXICO, Querétaro: J. N. Rose et al. 9720 (Holotype: US 00089577!) .

Quercus ocoteifolia Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 176 (1854) . Type:— MEXICO, Oaxaca: F. M. Liebmann 3520 (Lectotype: C 10012510! designated by Trelease (1924, p. 227, pl. 334a); isolectotypes: C!, CAS!, GH (as s.n.)!, UCSB (fragment)!).

Quercus orizabae Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 189 (1954) . Type:— MEXICO, Veracruz: F. M. Liebmann 3539 (Lectotype (designated here): C 10012521!; isolectotypes: C!, CAS!, GH (as s.n.)!, K (as s.n.)!, NY!, P (as s.n.)!, UC!, UCSB (fragment from C lectotype)!, UCSB (fragment)!). Trelease (1924) in pl. 233 shows two different specimens. In the caption to pl. 233 on p. 223 he states “The type specimens. (Mexico, Liebmann, 83-84=3539).” The upper image in pl. 233 is sheet 84 which we are designating as the lectotype, and the lower image in pl. 233 is sheet 83, an isolectotype also at C.

Quercus roseovenulosa Trel., Mem. Natl. Acad. Sci. 20: 182, pl. 366a (1924). Type:— MEXICO, México: C. A. Purpus 1797 (Holotype: MO-144176!; isotypes: F!, GH!, NY!, UC!, UCSB (fragment from MO holotype)!, US!) .

Quercus sideroxyla Bonpl. f. ciliifera Trel., Mem. Natl. Acad. Sci. 20: 170 (1924). Type:— MEXICO, Hidalgo: C. A. Ehrenberg 1279 (Lectotype (designated here): NY00023599 !; isolectotypes: BM!, P!, UCSB (fragment)!, VT!). Trelease (1924, p. 170) cited two different collections. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. laurina as broadly interpreted.

Quercus simillima Trel., Mem. Natl.Acad. Sci. 20: 180, pl. 363b (1924). Type:— MEXICO, San Luis Potosí: C. C. Parry & E. Palmer s. n. (1878) (Holotype: MO-143908!; isotype: UCSB (fragment from MO holotype)!).

Quercus tlapuxahuensis A.DC. var. obconica Trel., Mem. Natl. Acad. Sci. 20: 165, pl. 325b (1924). Type:— MEXICO, Ciudad de México: E. Bourgeau 1013 (Holotype: GH00055708!; isotypes: BM!, BR!, C!, G!, K!, LE!, MO!, MPU!, P!, UCSB (fragment)!, US!). Trelease (1924, p. 165, pl. 325b) considered “ Bourgeau, 1013 in part” at GH to be the type of Q. tlapuxahuensis var. obconica ; it is different from Bourgeau 1013 at G which is the holotype of Q. bourgaei .

Quercus tridens Bonpl., Pl. Aequinoct. 2: 56, pl. 96 (1809). Type :— MEXICO, Hidalgo: A. Bonpland 4061 (p.p.) (Holotype: P00669717 !; isotypes: P!, UCSB (fragment)!). There are two additional sheets of Bonpland 4061 (p. p.) at P that are isotypes of Q. tridens ( P00129746 ! and P0012974 !) .

Quercus xalapensis Bonpl. var. longifolia (Liebm.) Wenz., Jahrb. K ̂nigl. Bot. Gart. Berlin 3: 210 (1884).

Quercus zempoaltepecana Trel., Mem. Natl.Acad. Sci. 20: 164, pl. 323a (1924). Type :— MEXICO, Oaxaca: E. W. Nelson 672 (Holotype: US00089682!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. laurina as broadly interpreted.

Distribution: El Salvador, Guatemala, Honduras, Mexico (Aguascalientes, Chiapas, Ciudad de México, Coahuila,

Colima, Durango, Guanajuato, Guerrero, Hidalgo, Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo León ,

Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Tamaulipas, Tlaxcala, Veracruz, Zacatecas).

Quercus macdougallii Martínez,Anales Inst. Biol. Univ. Nac.Autón. México 34(1 & 2): 147 (1964). Type:— MEXICO, Oaxaca: T. B. MacDougall 503 (Holotype: MEXU00051895 !; isotypes: CAS!, GH!, ENCB (image)!, F!, MEXU!, MICH!, US!) .

Distribution: Mexico (Oaxaca).

Quercus magnoliifolia Neé, Anales Ci. Nat. 3: 268 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA23387 ! designated by Bartholomew & Almeda (2022, p. 29); isolectotype: MA!) .

Quercus chartacea Trel., Mem. Natl. Acad. Sci. 20: 61, pl. 59a (1924). Type:— MEXICO, Oaxaca: C. Seler & E. G. Seler 1631 (Holotype at B destroyed. Lectotype (designated here): NY01450026 !; isolectotype: GH!). The holotype was E. G. Seler 62, but no isotypes have been found. The lectotype is selected from the paratypes.

Quercus circinata Neé, Anales Ci. Nat. 3: 272 (1801). Type:— MEXICO, Guerrero: L. Neé s.n. (1791) (Lectotype: MA26472 ! designated by Muller & McVaugh (1972, p. 518); isolectotype: MA!) .

Quercus erubescens Trel., Repert. Spec. Nov. Regni Veg. 33: 315 (1934). Type:— MEXICO, Guerrero: L. Schultze 341 (Holotype: at B destroyed). The determination of Q. erubescens as probably Q. magnoliifolia is based on the Trelease (1934) description as no images or original specimens exist.

Quercus haematophlebia Trel., Mem. Natl. Acad. Sci. 20: 66, pl. 74 (1924). Type:— MEXICO, Nayarit: J. N. Rose 3363 (Holotype: US 00089527!) .

Quercus liebmannii Oerst. ex Trel., Mem. Natl. Acad. Sci. 20: 66, pl. 75a (1924). Type:— MEXICO, Oaxaca: F. M. Liebmann 3505 (Lectotype (designated here): C10012498 !; isolectotypes: C!, CAS!). Trelease (1924) on p. 67 wrote “ Liebmann, 89, 3505, 5742, May 1842, the type”, and on p. 67 in the caption to pl. 75a he wrote “The type (Mexico, Liebmann, 89=5742).” The specimen photographed in pl. 75a is, however, Liebmann 3505 not 5742. In fact, no specimen numbered 5742 has been found at C. Because of this ambiguity we are designating Liebmann 3505 at C as the lectotype. The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. magnoliifolia as broadly interpreted.

Quercus lutea Neé, Anales Ci. Nat. 3: 269 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA233384 ! designated by Bartholomew & Almeda (2022, p. 29); isolectotype: MA!) .

Quercus macrophylla Neé, Anales Ci. Nat. 3: 274 (1801). Type:— MEXICO, Guerrero: L. Neé s. n. (1791) (Lectotype: MA26466 ! designated by Muller & McVaugh (1972, p. 518); isolectotypes: MA!) .

Quercus magnoliifolia var. lutea (Neé) A.DC., Prodr. 16(2.1): 27 (1864).

Quercus magnoliifolia var. macrophylla (Neé) A.DC., Prodr. 16(2.1): 27 (1864).

Quercus poculifer Trel., Mem. Natl. Acad. Sci. 20: 64, pl. 69 (1924). Type:— MEXICO, Oaxaca: C. G. Pringle 4760 (Holotype: MO-144177!; isotypes: A!, BM!, BR!, CAS!, E!, ENCB (image)!, F!, G!, GH!, HBG (image)!, K!, KFTA (image)!, MEXU!, NDG (image)!, NY!, P!, PH!, UC!, UCSB (fragment)!, US!, VT!). The placement of this species needs additional study, but we consider the type to fall within the variation found within Q. magnoliifolia as broadly interpreted.

Quercus rubescens Trel., Repert. Spec. Nov. Regni Veg. 33: 316 (1934). Type:— MEXICO, Guerrero: L. Schultze 328 (Holotype: at B destroyed). The determination of Q. rubescens as Q. magnoliifolia is based on the Trelease (1934) description as no images or original specimens exist.

Quercus tepicana Trel., Mem. Natl. Acad. Sci. 20: 135, pl. 355 (1924). Type:— MEXICO, Nayarit: J. N. Rose s. n. (3 Aug 1897) (Holotype: US 00089679!) .

Distribution: Mexico (Aguascalientes, Baja California Sur, Chiapas, Ciudad de México, Coahuila, Colima, Durango,

Guanajuato, Guerrero, Hidalgo, Jalisco, México , Michoacán, Morelos, Nayarit, Oaxaca, Puebla, Querétaro, San Luis

Potosí, Sinaloa, Veracruz, Zacatecas).

Quercus mcvaughii Spellenb.,Amer. J. Bot. 79(10): 1200, fig. 1 (1992). Type :— MEXICO, Chihuahua: R. D. Spellenberg et al. 9821 (Holotype: NMC (accn. 059633) (image)!; isotypes: ASU (image)!, CAS!, CIIDIR (image)!, IBUG (image)!, MEXU!, NY!, MO!, US!, K!). This species is very close to Q. crassifolia and could be treated at the infraspecific level.

Distribution: Mexico (Chihuahua, Durango, Jalisco, San Luis Potosí, Sinaloa, Sonora).

Quercus meavei Valencia-A., Sabas & O. J.Soto, Phytotaxa 269(2): 121 (2016). Type:— MEXICO, Hidalgo: Valencia & Flores 5286b (Holotype: FCME; isotypes: HUMO, HGMO, HUAP). The holotype has not been seen. Some of the paratypes that have been seen are Q. xalapensis , but based on the protologue and illustrations, the type collection seems to represent a distinct species .

Distribution: Mexico (Hidalgo).

Quercus melissae Nixon & Barrie, Novon 25(4): 447, fig. 3 (2017). Type:— MEXICO, Chiapas: L. Kaplan & E. L. Davis 358 (p.p.) (Holotype: F!; isotypes: A!, F!, G!). There are also collections of Kaplan & Davis 358 at F which are Q. acutifolia so they are not isotypes. This species is close to Q. segoviensis , and additional study is needed to determine if it is distinct.

Distribution: Guatemala, Mexico (Chiapas).

Quercus mexicana Bonpl., Pl.Aequinoct. 2: 35, pl. 82 (1809). Type:— MEXICO, Hidalgo: A. Bonpland 4060 (Holotype: P00129716 !; isotypes: CGE (as s.n.)!, P!, UCSB (fragment)!).

Quercus castanea Neé var. integra Oerst., Bidra Kundsk. Egefamil. 362 (1871). Nom. nov. for Q. mexicana at the varietal level.

Quercus castanea var. mexicana (Bonpl.) A.DC., Prodr. 16(2.1): 72 (1864).

Quercus mexicana f. bonplandii Trel., Mem. Natl. Acad. Sci. 20: 174, pl. 347 (1924). Type :— MEXICO: A. Bonpland s. n. (Syntypes at B destroyed. Lectotype (designated here): P00754160 !) .

Quercus pablillensis C.H.Mull.,Amer. Midl. Naturalist 27(2): 477 (1942). Type:— MEXICO, Nuevo León: C. H. Muller & M. T. Muller 1100 (Holotype: A00055712 !; isotypes: BM!, F!, G!, P!, UCSB (fragment from A holotype)!, US!) .

Quercus rugulosa M.Martens & Galeotti, Bull. Acad. Roy. Sci. Bruxelles 10(1): 209 (1843). Type:— MEXICO, Hidalgo: H. Galeotti 116 (Holotype: BR0000006913618 !; isotypes: K!, P!, UCSB (fragment)!).

Quercus rugulosa f. subtruncata Trel., Mem. Natl. Acad. Sci. 20: 182 (1924). Type:— MEXICO, San Luis Potosí: E. Palmer 70 (Lectotype (designated here): A00969951 !; isolectotypes: BM!, F!, GH!, MEXU!, MO!, NY!, UC!, UCSB (fragment)!, US!). Trelease (1924, p. 182) cites two specimens without designating a type. The specimen at A has a determination in Trelease’s hand.

Distribution: Mexico (Chiapas, Ciudad de México, Coahuila, Guanajuato, Hidalgo, Jalisco, México, Michoacán,

Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Tamaulipas, Tlaxcala, Veracruz).

The occurrence of Q. mexicana in Jalisco, Michoacán, Nayarit, and Sinaloa reported by Villaseñor (2016) needs to be confirmed. The reports from Nayarit may possibly involve introgression with some other species.

Quercus microphylla Neé, Anales Ci. Nat. 3: 264 (1801). Type:— MEXICO, Guanajuato: L. Neé s. n. (1791) (Lectotype: MA26469! designated by Muller & McVaugh (1972, p. 519); isolectotypes: MA!, UCSB (fragment)!). Although there are differences between Q. microphylla , Q. cordifolia , and Q. striatula , they all form a broadly defined species. Further study would be important for this group.

Quercus cordifolia Trel., Mem. Natl. Acad. Sci. 20: 84, pl. 125b (1924). Type:— MEXICO, Coahuila: E. Palmer 1278 (Holotype: GH00055707 !; isotypes: AC!, K!, UCSB (fragment)!, US!) .

Quercus striatula Trel., Mem. Natl. Acad. Sci. 20: 93, pl. 151 (1924). Type:— MEXICO, Durango: R. Nelson 5001 (Holotype at B destroyed. Lectotype (designated here): GH00969979 !; isolectotypes: K!, US!). No isotypes of the original type, Endlich 3 (p.p.), have been found. The lectotype is selected from the paratypes.

Quercus striatula f. otinapensis Trel., Mem. Natl. Acad. Sci. 20: 94, pl. 152a (1924). Type:— MEXICO, Durango: E. Palmer 445 (Holotype: GH00055599 !; isotype: US!) .

Distribution: Mexico (Aguascalientes, Chihuahua, Ciudad de México, Coahuila, Durango, Guanajuato, Hidalgo,

Jalisco, México, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa, Tamaulipas, Tlaxcala ,

Veracruz, Zacatecas).

Quercus miquihuanensis Nixon & C.H.Mull., Brittonia 45(2): 152, fig. 3 (1993). Type:— MEXICO, Tamaulipas: K. C. Nixon et al. 4051-I (Holotype: BH (image)!). This species is part of the Q. hypoxantha complex of northeastern Mexico, but Q. miquihuanensis is sufficiently distinct to be treated here as a separate species.

Distributon: Mexico (Nuevo León, Tamaulipas) .

Quercus mohriana Buckley ex Rydb., Bull. New York Bot. Gard. 2(6): 219, pl. 31, figs. 1, 2 (1901). Type:— U.S.A., Texas: S. B. Buckley s. n. (1875) (Lectotype: NY00248752! designated by Trelease (1924 p. 220, pl. 153b) and redundantly relectotypificed by Dorr & Nixon (1985, p. 218); isolectotype: NY!). The distinction between Q. mohriana and Q. tinkhamii needs additional study. If Q. tinkhamii is included as a synonym, the distribution of Q. mohriana becomes much broader in Mexico.

Distribution: Mexico (Chihuahua, Coahuila, Nuevo León), U.S.A. (New Mexico, Oklahoma, Texas).

Quercus muehlenbergii Engelm., Trans. Acad. Sci. St. Louis 3: 391 (1877). Type:— U.S.A.: F. T. Pursh s. n. (1806) (Holotype: PH 00038773 (image)!).

Quercus acuminata (Michaux) Sarg. non Roxb. (1832), Silva 8: 55 (1895).

Quercus brayi Small, Bull. Torrey Bot. Club 28(6): 358 (1901). Type :— U.S.A., Texas: H. Lacey s. n. (1899–1900) (Holotype: NY00248621 !; isotype: CAS!) .

Quercus muehlenbergii var. brayi (Small) Sarg., Bot. Gaz. 65(5): 442 (1918).

Quercus sentenelensis C.H.Mull., Amer. Midl. Naturalist 18(5): 849 (1937). Type:— MEXICO, Coahuila: F. L. Wynd & C. H. Muller 629 (Lectotype (designated here): ILL00007587 !; isolectotypes: A!, ILL!, K!, MO!, NY!, UCSB (fragment)!). The two specimens at ILL were marked as co-types.

Distribution: Canada (Ontario), Mexico (Chihuahua, Coahuila, Nuevo León, Sonora, Tamaulipas), U.S.A. (Alabama,

Arkansas,Connecticut,Florida,Georgia,Illinois,Indiana,Iowa,Kansas,Kentucky,Louisiana,Maryland,Massachusetts,

Michigan, Minnesota, Mississippi, Missouri, Nebraska, New Jersey, New Mexico, New York, North Carolina, Ohio,

Oklahoma, Pennsylvania, South Carolina, Tennessee, Texas, Vermont, Virginia, West Virginia, Wisconsin).

Quercus nanchititlensis B.M.Barthol. & Almeda , nom. nov. here designated for Q. platyphylla W.F. Warb. (1939) non Morogues (1877). Type:— MEXICO, México: G. B. Hinton 6360 (Holotype: K000575020 ! and K000575021 !; isotypes: A!, BM!, F!, G!, K!, NY!). The holotype is mounted on two sheets with the second sheet consisting of additional leaves that would not fit on the first sheet.

Quercus platyphylla E.F.Warb. non Morogues(1877), Bull.Misc. Inform.Kew1939(2):85 (1939).See Q. nanchititlensis above.

Distribution: Mexico (Guerrero, Jalisco, México, Michoacán).

Quercus nudinervis Liebm., Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 182 (1854) . Type:— MEXICO, Sinaloa: B. Seemann 1457 (Holotype: K000512617 !; isotypes: BM!, GH (as s.n.)!, K!, UCSB (fragment)!). The holotype at K has the determination in Liebmann’s hand.

Distribution: Mexico (Durango, Jalisco, Nayarit, Sinaloa).

Quercus oblongifolia Torr., Rep. Exped. Zuni & Colorado Rivers 173, pl. 19 (1853). Type:— U.S.A., New Mexico: D. Woodhouse s. n. (Holotype: NY00253711 !; isotype: MO (fragment)!).

Quercus perpallida Trel., Mem. Natl. Acad. Sci. 20: 91, pl. 147 (1924). Type :— MEXICO, Sonora: J. N. Rose et al. 13089 (Holotype: US 00089592!; isotype: NY!) .

Distribution: Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Nuevo León, Sinaloa,

Sonora, Tamaulipas, Zacatecas), U.S.A. (Arizona, New Mexico, Texas) .

Quercus obtusata Bonpl., Pl. Aequinoct. 2: 26, pl. 76 (1809). Type:— MEXICO, Michoacán: A. Bonpland 4329 (Holotype: P00129713 !; isotypes: P!, UCSB (fragment from P holotype)!, UCSB (fragment)!).

Quercus atriglans E.F.Warb., Bull. Misc. Inform. Kew 1939 (2): 88 (1939). Type:— MEXICO, México: G. B. Hinton 6549 (Holotype: K000512995 !; isotypes: BM!, F!, G!, GH!, K!, MO!, UCSB (fragment)!, VT!).

Quercus centralis Trel. f. heterophylla Trel., Mem. Natl. Acad. Sci. 20: 62 (1924). Type :— MEXICO, Puebla: G. Arsène 7052 (Holotype at B destroyed. Lectotype (designated here): US 00089460!; isolectotypes: MO!, MPU!) .

Quercus crenatifolia Trel., Mem. Natl. Acad. Sci. 20: 57, pl. 52b (1924). Type:— MEXICO, Jalisco: M. E. Jones 440 (Holotype: MO-143884!; isotypes: DS!, UCSB (fragment)!, US!) .

Quercus innuncupata Trel., Mem. Natl. Acad. Sci. 20: 77, pl. 108 (1924). Type:— MEXICO, Michoacán: G. Arsène 5826 (Holotype: F (accn. 416692)!; isotypes: ILL!, MPU!, PH!, US!) .

Quercus obtusata var. pandurata (Bonpl.) A.DC., Prodr. 16(2.1): 27 (1864).

Quercus pandurata Bonpl., Pl. Aequinoct. 2: 28, pl. 77 (1809). Type:— MEXICO, Michoacán: A. Bonpland 4330 (Holotype: P00129711 !; isotypes: B-W!, P!, USSB (fragment)!).

Quercus panduriformis Trel., Mem. Natl. Acad. Sci. 20: 62, pl. 62b (1924). Type:— MEXICO, Colima: H. Ross 519 (Lectotype (designated here): M-0244466!). Trelease (1924, p. 63) cites Endlich 1357 as the type of Q. panduriformis which was undoubtedly at B and destroyed, but in the caption to pl. 62b he also cites Ross 579 at M as the type. No specimen of Ross 579 was found at M, but there is a specimen of Ross 519 which is the specimen photographed in pl. 62b. Trelease misread the “1” as a “7” which is an easy mistake to make. We have determined the M specimen of Ross 519 to be Q. obtusata . Note that Trelease (1924, p. 62) in the specimens examined statement for Q. obtusata does cite “? Ross, 519 ” indicating that he was uncertain, at least at one time, if Ross 519 was actually Q. obtusata , but he decided that it belonged to his new species Q. panduriformis . Because two different collections were cited by Trelease as the type of Q. panduriformis , one of which is now lost, as well as a typographic error in citing the extant specimen, we are designating Ross 519 as the lectotype of Q. panduriformis .

Quercus panduriformis f. rubrinervis Trel., Mem. Natl. Acad. Sci. 20: 63 (1924). Type:— MEXICO, Guerrero: E. Langlassé 600 (Lectotype (designated here): US00089600!; isolectotypes: G!, GH!, K!, MPU!, P!). Trelease (1924, p. 63) cites two Langlassé collections without designating one as the type.

Quercus reticulata Bonpl. subsp. rhodophlebia (Trel.) A.Camus f. crenata (Trel.) A.Camus, Chênes Texte 2: 581 (1939).

Quercus rhodophlebia Trel. f. crenata Trel., Mem. Natl. Acad. Sci. 20: 74 (1924). Type:— MEXICO, Michoacán: G. Arsène 6018 (Lectotype (designated here): ILL00007616 !; isolectotypes: A!, CAS!, MO!, MPU!, UCSB (fragment)!, US!). Trelease (1924, p. 74) cited two different Arsène collections without indicating where they were deposited.

Distribution: Mexico (Aguascalientes, Chiapas, Ciudad de México, Colima, Durango, Guanajuato, Guerrero, Hidalgo,

Jalisco, México, Michoacán, Morelos, Nayarit, Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Sinaloa ,

Tamaulipas, Tlaxcala, Veracruz, Zacatecas).

Quercus oleoides Schltdl. & Cham., Linnaea 5(1): 79 (1830). Type :— MEXICO, Veracruz: C. J. W. Schiede 23 (Holotype at B destroyed. Lectotype (designated here): HAL0060354 !; isolectotypes: HAL!, LE!) .

Quercus lutescens M.Martens&Galeotti,Bull.Acad.Roy.Sci.Bruxelles 10(1): 219(1843). Type:— MEXICO, Veracruz: H. Galeotti 91 (Lectotype: BR0000006915636 ! designated by Trelease (1924, p. 222, pl. 194); isolectotypes: BR!, G!, K!, P!, UCSB (fragment)!). Although M. Martens & Galeotti (1843) cited only one collection number, there are two sheets of this number at BR marked as types, one of which was designated by Trelease as “The type.”

Quercus oleoides var. australis Trel., Mem. Natl. Acad. Sci. 20: 114, pl. 192b, 193 (1924). Type:— COSTA RICA, Heredia: H. Pittier 2607 (Holotype: BR0000006911669 !; isotypes: US!, UCSB (fragment from BR holotype)!, UCSB (fragment)!).

Quercus oleoides f. lutescens (M.Martens & Galeotti) Trel., Mem. Natl. Acad. Sci. 20: 114, pl. 194 (1924).

Quercus oleoides var. lutescens (M. Martens & Galeotti) A.Camus, Chênes Atlas 2: 64 (1935).

Quercus retusa Liebm. non Raf. (1838), Overs. Kongel. Danske Vidensk. Selsk. Forh. Medlemmers Arbeider 1854: 187 (1854). Type:— COSTA RICA, Heredia: A. Oersted 4 (Holotype: C 10012516!; isotype UCSB (fragment from C holotype)!).

Distribution: Belize, Costa Rica, Guatemala, Honduras, Mexico (Chiapas, Ciudad de México, Guerrero, Hidalgo,

Nuevo León, Oaxaca, Puebla, Querétaro, San Luis Potosí, Tabasco, Tamaulipas, Veracruz), Nicaragua.