Anomalomorpha pilbara Hutchinson & Allsopp, 2021

Hutchinson, Paul M. & Allsopp, Peter G., 2021, Revision of Anomalomorpha Arrow, 1908 and Enracius Dechambre, 1999, with Erbmahcedius new genus from southeastern Australia (Coleoptera: Scarabaeidae: Dynastinae: Pentodontini: Cheiroplatina), Zootaxa 5072 (5), pp. 439-462 : 450-456

Anomalomorpha pilbara Hutchinson & Allsopp , new species urn:lsid:zoobank.org:act:5303AD6F-18BF-494B-A592-0B386118C436

Type series. Holotype male. AUSTRALIA, WESTERN AUSTRALIA: “ West Angelas at 23.06°S, 118.46°E, 29 June 1984, R.P. McMillan | in litter | Anomalomorpha ” | aedeagus on card; in WAM.

Allotype. AUSTRALIA, WESTERN AUSTRALIA: 1♀: same data as holotype; in WAM.

Description of holotype male ( Figs. 28–34 , 42 , 46–47 ). Left mesotarsomeres 4–5, right mesotarsomeres 5, left metatarsomere 5, right metatarsomeres 4–5, left and right maxillary palpomere 4 missing, only antennal scape and pedicel present. Body 16.6 mm long, pronotum 8.4 mm wide at widest point, elytra 10.5 mm wide; form convex depressed; dorsal surface matt, head, pronotum, elytra and pygidium black; scutellum, elytral suture, margin, disc, sternum, abdominal ventrites and legs dark brown to black. Maxillary palp with palpomeres 1 and 3 subequal in length; mandibles with lateral margin bearing long setae; mentum wide, bearing very long setae laterally, sparsely setose medially. Antenna with scape bearing long setae, pedicel with sparse short setae. Clypeus transverse, almost trapezoidal, distance between outer edges of canthi 2.0 times interocular distance, anterior angles rounded, anteriorly emarginate, truncate face vertical with shallow punctures, glabrous, dorsal surface rugulose; clypeofrontal ridge posteriorly angulate, weakly raised leading to median node. Frons rugulose; ocular canthi angulate to line of clypeus then continuously arcuate, broad, dorsal surface rugulose, anterior margin bearing row of long stout setae. Pronotum widest post midlength, divergent from base and arcuate to acute anterior angles, recurved to linear transverse anterior margin, anterior margin membranous; lateral and anterior ridges complete, obsolete across base; disc evenly convex, bearing sparse micropunctures becoming larger and denser adjacent to lateral margins. Scutellum sparsely punctate on disc. Elytral striae linear-punctate, intervals impunctate, lateral and apical declivity rugulose, suture punctate-striate, epipleura non-emarginate, continuous to pygidium, lateral setae sparse and ending adjacent to pygidium. Pygidium moderately convex, disc sparsely micropunctate, denser at lateral margins where bearing sparse, long setae, apical ridge glabrous. Prosternal process long, slender, apex angled posteriorly and setose. Metasternum and metepisternum densely punctate, laterally rugulose and bearing long, lightbrown setae. Abdominal sternites with a line of continuous setae across midlength, sternite 7 apical margin bearing row of sparse, long setae. Protibia tridentate, denticles equidistant; meso- and metatibia bicarinate, apical cilia long, sharp, with interspersed shorter cilia; metafemur broad, anterior margin convex, posterior margin linear; metatibia stout, flat on inner surface with dorsal margin concave post medial carina, inner spur 1.5 times length of outer spur, both narrow, straight and tapering to acute apex; metatarsomeres 1–3 combined about 0.8 times length of metatibia. Metacoxal posterior margin angulate at midlength, then linear to narrow arcuate posterolateral angle. Genitalia (in posterior view) with parameres laterally along basal half shallowly concave, midlength with low, rounded, obtusely angled process, apices tapering to acute apex.

Description of allotype female ( Figs. 35–40 ). Left protarsomeres 2–5, right protarsomeres 3–5, left mesotarsomeres 3–5, right meso-leg, metatarsomeres 3–5, right labial and maxillary palps missing, only antennal scape and pedicel present. Body 17.4 mm long, pronotum 8.1 mm wide at widest point, elytra 10.1 mm wide; form convex depressed; dorsal surface matt, head, pronotum, elytra and scutellum black; elytral suture, margins, sternum, abdominal ventrites, pygidium and legs brown to dark brown. Maxillary palpomeres 1 and 3 subequal in length, palpomere 2 longer but shorter than palpomere 4, palpomere 4 about 3 times longer than greatest width and bearing trace of narrow, elongate sensorium on basal two-thirds; mandibles with lateral margin bearing long setae; mentum wide, bearing very long setae laterally, sparsely setose medially (most abraded). Antennal scape bearing long setae, pedicel with sparse, short setae. Clypeus transverse, almost trapezoidal, distance between outer edges of canthi 1.9 times interocular distance, anterior angles rounded, anteriorly emarginate, truncate face vertical with shallow punctures, glabrous, dorsal surface rugulose; clypeofrontal ridge posteriorly angulate, weakly raised leading to medial node. Frons rugulose; ocular canthi angulate to line of clypeus then continuously arcuate, broad, dorsal surface rugulose, anterior margin bearing row of long, stout setae (most abraded). Pronotum widest post midlength, divergent from base and evenly arcuate to acute anterior angles (almost right-angle), recurved to linear transverse anterior margin, anterior margin membranous laterally; lateral and anterior ridges complete, obsolete across base; disc evenly convex, bearing sparse micropunctures becoming larger and denser adjacent to lateral margins. Scutellum sparsely punctate on disc. Elytral striae linear-punctate, intervals impunctate, lateral and apical declivity rugulose, suture punctate-striate, epipleura non-emarginate, continuous to pygidium, lateral setae sparse, ending adjacent to pygidium. Pygidium moderately flat, disc sparsely micropunctate, denser at lateral margins where bearing sparse, long setae, apical ridge glabrous. Prosternal process long, slender, apex angled posteriorly and setose. Metasternum and metepisternum densely punctate, laterally rugulose and bearing long, light-brown setae. Abdominal sternites with a line of continuous setae adjacent to posterior margin, sternite 6 posteriorly with membranous, transverse groove, sternite 7 apical margin bearing row of sparse, long setae. Protibia tridentate, denticles non-equidistant with basal denticle closer to medial than medial is to apical; meso- and metatibia bicarinate, apical cilia long, sharp with interspersed shorter cilia; metafemur broad, anterior margin convex, posterior margin linear; metatibia stout, flat on inner surface with dorsal margin concave post medial carina, inner spur 1.5 times length of outer spur, both moderately broad, widest preapically, inner spur straight, outer spur slightly curved; metatarsomeres 1–2 combined about 0.5 times length metatibia. Metacoxal posterior margin sublinear to narrow arcuate posterolateral angle.

Sexual differential diagnosis. Female differs from male by the following combination of characters: protibial denticles non-equidistant, metatibial spurs broader, metacoxa with posterior margin sublinear; abdominal sternites with setae along posterior margin, pygidium quite flat.

Differential diagnosis of males (female A. monteithi unknown). Given the paucity and condition of specimens, we did not attempt cleaning. Anomalomorpha pilbara can be distinguished from its congeners by the following combination of characters:

• Ocular canthus with lateral margin continuously arcuate (weakly arcuate then posteriorly angulate in A. anthracina and A. monteithi );

• Distance between outer edges of canthi 1.9 times interocular distance (1.8–1.9 times in A. anthracina and 1.6- 1.65 times in A. monteithi );

• Pygidium with apical ridge glabrous (setose in A. anthracina and A. monteithi );

• Metacoxa with posterior margin angulate at midlength, then narrowly arcuate at posterolateral angle (broadly arcuate from midlength through posterolateral angle in A. anthracina ; sublinear then arcuate through posterolateral angle in A. monteithi );

• Metafemur with posterior margin linear (weakly concave in A. anthracina , sublinear in A. monteithi );

• Parameres with lateral tooth obtusely rounded (angulate and prominent in A. anthracina and A. monteithi );

• Parameres (posterior view) tapering to acute apices that are curved inwards (parallel to truncate apices in A. anthracina ; tapering to acute apices in A. monteithi ; apices of both curved slightly outwards).

Etymology. Named for the area of the type locality that is noted for its red earth, as attached to the holotype; a noun in apposition. The Pilbara Aboriginal Language Centre Wangka Maya says that the name for the Pilbara region derives from the word bilybara, meaning “dry” in the Nyamal and Banyjima languages ( Anonymous 2021).

Distribution ( Fig. 51 ). The type locality is in an area classified as Köppen-Geiger climate BSh (hot semi-arid) on the edge of a larger area of BWh (hot desert). It has been modified by mining, but the surrounding area ( Fig. 50 ) is red, rocky ranges with multiple gorges and creek lines and covered with Eucalyptus / Corymbia (Myrtaceae) with an understorey predominately of Acacia over spinifex ( Triodia ; Poaceae ) and buffel grass ( Cenchrus ciliaris Linnaeus ; Poaceae ).

Figures

FIGURE 50. Mount Robinson Gorge, about 20 km NNE of West Angelas and with similar topography and vegetation to that originally at the type locality of Anomalomorpha pilbara Hutchinson & Allsopp new species.

FIGURE 51. Known distribution of Anomalomorpha species and Erbmahcedius giveni (Carne, 1957).

FIGURES 28–30. Anomalomorpha pilbara Hutchinson & Allsopp new species, holotype male. 28, Dorsal view; 29, ventral view; 30, lateral view.

FIGURES 31–34. Anomalomorpha pilbara Hutchinson & Allsopp new species, holotype male. 31, Head dorsal view; 32, head lateral view; 33, pygidium; 34, labels and mounted aedeagus.

FIGURES 41–43. Anomalomorpha metacoxae. 41, Anomalomorpha anthracina Arrow, 1908; 42, A. pilbara Hutchinson & Allsopp new species; 43, A. monteithi Allsopp, 2000.

FIGURES 44–49. Anomalomorpha male aedeagi. 44–45, Anomalomorpha anthracina Arrow, 1908. 44, parameres dorsal; 45, aedeagus lateral; 46–47, A. pilbara Hutchinson & Allsopp new species. 46, parameres dorsal; 47, aedeagus lateral; 48–49, A. monteithi Allsopp, 2000. 48, parameres dorsal; 49, aedeagus lateral. Arrows show regions of major differences.

FIGURES 35–37. Anomalomorpha pilbara Hutchinson & Allsopp new species, allotype female. 35, Dorsal view; 36, ventral view; 37, lateral view.

FIGURES 38–40. Anomalomorpha pilbara Hutchinson & Allsopp new species, allotype female. 38, Head dorsal view; 39, head lateral view; 40, pygidium.