Tonnoiriella rhodesica, Wagner & Withers, 2020

Wagner, Rüdiger & Withers, Phil, 2020, The West-Palearctic species of the genus Tonnoiriella Vaillant, 1971 (Diptera: Psychodidae, Psychodinae), Zootaxa 4728 (2), pp. 183-210 : 195-196

Tonnoiriella rhodesica sp. nov.

( Fig. 23 )

Material: 1♂, holotype, Greece, Rhodes , Ixos, 24 May 1960, leg. Jung ( RW) .

Description: Head with eyebridge of 3 facet rows; distance between eyes 3.7 facet diameters; a single row of postocular bristles. Antenna with scape, pedicel and 14 flagellomeres: absolute length 0.042-0.048-0.036-0.038- 0.042-0.042-0.042-0.038-0.036-0.032-0.032-0.032-0.03-0.022- 0.022-0.026 mm; relative length 21-24-18-19-21- 21-21-19-18-16- 16-16-15-11 -11-13; flagellomeres 5–11 with a pair of digitiform curved ascoids in the distal third. Palpus segments, absolute length: 0.06-0.082- 0.09-0.164 mm; relative lenght 30-41-45-82.

Wing 2.75 times longer than wide; length 1.74 mm, width 0.63 mm.

Male terminalia ( Fig. 23 ) with hypandrium broadened along almost the entire ventral side. Gonocoxites approximately 3x longer than wide, straight; inner side with basal broadening with a group of 10 longer setae, just before middle with a single long seta; gonostylus curved, shorter than gonocoxite.

Parameres with a broad ventral bridge, lobes do not reach the basis of the long seta; ventrally in the midst with a racket-shaped brownish mark of the aedeagus sheath; above a pair of processes as long as the more ventral processes that do not meet in the middle; further dorsal large rhomboid condyles that form the median keel.

Basiphallus 3x longer than wide, bilobed; ventral lobe bifid, ventral part connected to the distiphallus, longer part slightly bent with blunt tip; distiphallus with both ends rounded, the broader end with a short basal projection, the smaller end rounded wider at tip. The dorsal lobe of the basiphallus distally fixed to an anchor shaped cross distiphallus sclerite as broad as the ventral distiphallus.

Epandrium rhomboid, epandrial processes almost straight, apically with 10 feathered tenacula.

Distribution: Island of Rhodes ( Greece).

Etymology: the epithet is derived from Rhodes island, where the species was collected.

Remarks: size and shape of the hypandrium and the dorsal distiphallus indicate that the species is part of the T. sieberti -group; these features distinguish the new species form its congeners. The very wide and broad hypandrium has the effect that the gonocoxites are situated more lateral and run almost parallel.

Figures

FIGURES 15–25: 15 T. sieberti genitalia ventral view; 16 T. ikariae genitalia ventral view; 17–21 T. italiae, genitalia ventral view of specimen from northern Italy (17), Sicily (18), aedeagus of specimen from N Italy (19), Sicily, Villa doso (20) and Sicily Castelbuono (21); 22 T. turcica genitalia ventral view; 23 T. rhodesica genitalia ventral view; 24 T syriensis genitalia ventral view, 25 T. syriensis epandrium and epandrial processes oblique view.