Amplaria umatilla, Shear, 2021

Shear, William A., 2021, The millipede family Striariidae Bollman, 1893. II. New records and species of the genus Amplaria Chamberlin, 1941 (Diplopoda, Chordeumatida, Striarioidea), Zootaxa 4908 (2), pp. 205-224 : 218

Amplaria umatilla , new species

Figs. 35–37 , 54 , 55

Types: Male holotype and male paratype from Tiger Creek , Umatilla National Forest ,

45.9800500, -118.0522833 °, 2417’ asl, Umatilla Co., Oregon, collected 24 October 2003 by W. Leonard; additional male paratype from same locality, but 45°58.685’, -118°03.137’. Parts of these specimens are mounted on SEM stub WAS34-11 .

Diagnosis: Distinct in the form of the gonopods ( Figs. 37 , 54 , 55 ), and in the angular, overlapping median termini of the pleurotergal bars of ring 7 ( Fig. 35 ).

Etymology: The species name, a noun in apposition, is from the Umatilla National Forest and Umatilla County, Oregon.

Description: Male holotype. Length, about 11 mm, width 1.0 mm. Five ommatidia in two rows. Labrum with long, slightly sinuous hooks at each angle. Mandibles not modified. Legpairs 1 and 2 similar in size ( Fig. 35 ); legpair 1 without needle-like long setae, tarsi with spatulate, twisted setae ventrally. Second trochanters with moderate posteriorventral processes, bearing groups of specialized setae. Legs 3 with coxal flasks long, asetose, diverging; prefemora distally expanded, telopodites not strikingly reduced ( Fig. 35 ). Pleurotergal bars of ring 3 expanded, angular distally, overlapping in midline. Crests moderate. Pygidium short. Color uniform medium brown.

Gonopod ( Figs. 37 , 54 , 55 ) anterior angiocoxites with broad, two-toothed lateral process, smaller teeth distally on posterior surface. Posterior angiocoxites narrow at origin, broadening distally, sheathing 3 or possibly 4 long flagellocoxites. Ninth legpair with small distal coxal process, flattened telopodites articulating loosely with depression on pleural margin of ring 7 ( Fig. 36 ).

Females not collected.

Distribution: Known from the two close-by localities listed under Types.

Figures

FIGS. 32–36. Amplaria staceyi, A. umatilla. Figs. 32–34. Amplaria staceyi. Fig. 32. Apophysis of left leg 2 of male, ventral view. Fig. 33. Left gonopod of male, lateral view. Fig. 34. Right gonopod of male, mesal view. Figs. 35, 36. A. umatilla. Fig. 35. Second and third legpair of male, ventral view. Fig. 36. Gonopod complex, lateral view.

FIGS. 37–40. Amplaria umatilla, A. cervus. Figs. 37. Amplaria umatilla, right gonopod, mesal view. Figs. 38–40. A. cervus. Fig. 38. Right gonopod, mesal view. Fig. 39. Legpair 9 of male, anterior view. Fig. 40. Right second coxa and trochanter, posterior view.

FIGS. 45–54. Drawings of gonopods of Amplaria species. Fig. 45. Left gonopod of A. nazinta, lateral view. Fig. 46. Tip of anterior angiocoxite of A. nazinta gonopod, lateral view. Fig. 47. Right gonopod of A. crawfordi, mesal view. Fig. 48. Tip of anterior angiocoxite of A. crawfordi, lateral view. Fig. 49. Right gonopod of A. fontinalis, mesal view. FIG. 50. Tip of anterior angiocoxite of A. fontalis, posteriolateral view. Fig. 51. Right gonopod of A. rykkenae, mesal view. Fig. 52. Right gonopod of A. rykkenae, lateral view. Fig. 53. Right gonopod of A. baughi, lateral view. Fig. 54. Right gonopod of A. umatilla, lateral view.

FIGS. 55–59. Drawings of gonopods of Amplaria species. Fig. 55. Right gonopod of A. umatilla, mesal view. Fig. 56. Right gonopod of A. cervus, mesal view. Fig. 57. Right gonopod of A. mendocino, mesal view. Fig. 58. Right gonopod of A. flucticulus, lateral view. Fig. 59. Right gonopod of A. flucticulus, mesal view.