Phasgonophora granulis Delvare, 2020

Gul, Muhammad Athar, Soliman, Ahmed Mostafa, Gadallah, Neveen Samy, Al Dhafer, Hathal Mohammed & Delvare, Gerard, 2020, The genus Phasgonophora Westwood, 1832 (Hymenoptera, Chalcididae) in Saudi Arabia: re-evaluation of its limits and description of three new species, Journal of Hymenoptera Research 76, pp. 1-38 : 1

publication ID	https://dx.doi.org/10.3897/jhr.76.38340
publication LSID	lsid:zoobank.org:pub:8C7E1DDE-BCFA-47C0-A38D-18458AD9221E
persistent identifier	https://treatment.plazi.org/id/5EB72879-1E9C-4A89-BCD8-FF37534B7172
taxon LSID	lsid:zoobank.org:act:5EB72879-1E9C-4A89-BCD8-FF37534B7172
treatment provided by	Journal of Hymenoptera Research by Pensoft
scientific name	Phasgonophora granulis Delvare
status	sp. nov.

Treatment

Phasgonophora granulis Delvare sp. nov.

Figs 7A-E View Figure 7 , 8A-D View Figure 8 , 9A, B View Figure 9

Type material.

Holotype ♀: Kingdom of Saudi Arabia, Al-Baha, Al Mikhwa (Shada Al-Ala Natural Reserve) [19°50'51"N, 41°18'06.12"E, Alt. 1358 m], 14.IV.2016, e.l. Acacia , leg. D. Baiocchi [KSMA]. Paratypes: 7♀, same data as holotype [KSMA]; 2♀, same data as holotype [BMNH]; 2♀, same data as holotype [USNM]; 3♀, same data as holotype [EFC]; 2♀, same data as holotype but differing as for the coordinates [19°51'39.96"N, 41°18'15.84"E, Alt. 1248 m] and collection date, 29.III.2017 [KSMA]; 2♀, Kingdom of Saudi Arabia, Asir, Muhayil, Wadi Sabian (28 km SSE of Muhayil) [18°17'54.89"N, 42°07'41.11"E, Alt. 809 m], 05.IV.2017, e.l. Acacia , leg. D. Baiocchi [KSMA].

Diagnosis.

Gaster longer than mesosoma and acuminate, with syntergum longer than mesotibia (1.15 ×) (Fig. 9A, B View Figure 9 ); gena densely and entirely punctured (Figs 7D View Figure 7 , 8A View Figure 8 ); occiput completely strigulose (Fig. 7A View Figure 7 ); flagellum filiform, with all flagellomeres much longer than wide, F1 2.5 × as long as wide (Fig. 7C View Figure 7 ); mesosomal dorsum somewhat flattened (Fig. 8A View Figure 8 ); pronotal collum and mesonotum cristate punctured (Fig. 8A View Figure 8 ); axillae densely setose, setation masking integument posteriorly (Fig. 7E View Figure 7 ); propodeum with sharp spiracular teeth (Fig. 7E View Figure 7 ); mesepisternum with epicnemial carina forming sharp tooth mesoventrally (Fig. 8A View Figure 8 ); fore wing with dense but short setation, and pigmented track of Rs and r-m (Fig. 8D View Figure 8 ); Gt1 with weak wrinkles dorsally (Fig. 9A View Figure 9 ); Gt6 with deep punctures and very small, hardly visible, spiracle; cerci removed from base of syntergum, situated at half of its length (Fig. 9A, B View Figure 9 ).

Etymology.

The name is chosen in reference to the secondary sculpture of the areoles on the head and mesonotum, giving to them a dull, granulose appearance (see Fig. 8B View Figure 8 ).

Condition of holotype.

Specimen glued on rectangular card. Head and mesosoma partly covered with a thin layer on the bottom of areoles; second to fifth tergites with sides wide apart from each other, probably resulting from immersion in some medium.

Description of female holotype.

Body 8.4 mm. Colour. Head and mesosoma entirely black (Fig. 7A-E View Figure 7 ), metasoma brown (Fig. 9A View Figure 9 ), with syntergum darker laterally (Fig. 9B View Figure 9 ); tegula brownish (Fig. 8A View Figure 8 ); fore and mid legs dark brown, but knees, apex of tibiae and tarsi testaceous; hind leg dark brown (Fig. 8C View Figure 8 ), ventral femoral teeth and ventral side of tibia black (Fig. 8C View Figure 8 ), tarsus lighter; antenna entirely black (Fig. 7C View Figure 7 ); wings hyaline, veins dark brown (Fig. 8D View Figure 8 ).

Head (Figs 7A, B, D View Figure 7 , 8A View Figure 8 ). Hardly wider than mesosoma; with moderately dense setation, the setae long, thin and suberect, regularly distributed according to punctures; lower face and frons strongly convex, without preorbital ridges (Fig. 7D View Figure 7 ); vertex, frons and lower face densely punctured (Fig. 7B, D View Figure 7 ), gena more coarsely punctured (Fig. 8A View Figure 8 ); both mandibles 3-teethed (Fig. 7D View Figure 7 ), lower tooth the largest and somewhat removed from the mid one; clypeus hardly protruding at free margin (Fig. 7D View Figure 7 ); edge of oral fossa thickened (Fig. 7D View Figure 7 ); tentorial pits absent (Fig. 7B View Figure 7 ); scrobal depression entirely transversely strigose, reaching median ocellus (Fig. 7B View Figure 7 ); interantennal projection (Fig. 7B View Figure 7 ) strongly compressed laterally, narrower than antennal torulus, punctulate on front surface (one row of punctures only), narrowly produced upwards, but without flange above it; vertex with short but distinct carina behind median ocellus (Fig. 7A View Figure 7 ); POL 6 × OOL (Fig. 7A View Figure 7 ); occiput entirely strigulose, except for a row of punctures behind posterior edge of eye (Fig. 7A View Figure 7 ).

Antenna (Fig. 7C View Figure 7 ). Scape linear, its apex with level of vertex; pedicel 1.2 × as long as wide, with slight basal bottle neck; anellus slightly transverse, 0.8 × as long as wide, tapering basally; funicular segments pubescent, bearing numerous, not raised multiporous plate sensilla in several intricate rows; F1 2.2 × as long as wide, shorter than F2; F2 as long as F3; F7 1.64 × as long as wide; clava 2-segmented (suture nevertheless hardly distinct), narrowly rounded apically.

Mesosoma (Figs 7E View Figure 7 , 8A View Figure 8 ). With setae about twice as long puncture diameter, curved and suberect; setae regularly distributed on punctures, but pronotum in front of prepectus, axillae and propodeum laterally around the spiracle, densely setose, the setae adpressed there; dorsum of mesosoma somewhat flattened (Fig. 8A View Figure 8 ), with dorsal outline of pronotal collar and mesonotum straight; punctures with secondary, very fine, sculpture on their bottom (visible only at very high magnification: 800 ×) (as in Fig. 8B View Figure 8 ), thus appearing dull; pronotal collum transversely strigose (Fig. 7E View Figure 7 ); pronotal collar and mesonotum uniformly cristate punctured, the anterior wall of punctures forming crests (Fig. 8A View Figure 8 ); pronotal collar with shallow mesal depression, its sides strongly convex (Fig. 8A View Figure 8 ); pronotal carina visible laterally, forming a tooth in dorsal view; lateral panel mostly flat, with longitudinal carinulae dorsally and raised curved carina ventrally; notauli impressed (Fig. 7E View Figure 7 ); tegula with a tuft of about 10 setae anteriorly (Fig. 8A View Figure 8 ); mesoscutellum truncate anteriorly, rounded apically, its posterior margin raised and surpassing postscutellum (Fig. 7E View Figure 7 ); propodeum strongly sloping anteriorly, more strongly so posteriorly, with sharp spiracular tooth (Fig. 7E View Figure 7 ) and two irregular costulae; mesepisternum with mesodiscrimen as faint carina all over, without ventral fovea; epicnemial carina moderately raised laterally, strongly protruding mesoventrally, appearing as a sharp tooth in lateral view (Fig. 8A View Figure 8 ); ventral shelf in mesepisternum very weakly sculptured; adscrobal area, mesepimeron, and metepimeron coarsely areolate, the later bearing long setae; femoral depression of mesepisternum with only a few low carinae (Fig. 8A View Figure 8 ).

Wings (Fig. 8D View Figure 8 ). Fore wing densely setose but bare on basal cell, basal and cubital folds, marginal cell with a single, incomplete row of setae on the underside; setae generally very short on the disc of the wing, somewhat longer below MGV, PMV and Rs track; setae uniformly short on the underside of wing; MGV 0.27 × as long as SMV; PMV 0.38 × as long as MGV; STV 0.33 × as long as PMV; hind wing with 4 hamuli, the basal one the largest, removed from the followings.

Legs (Fig. 8C View Figure 8 ). Procoxa with deep depression anteriorly, margined dorsolaterally with carina raised into flange. Protibia with sharp, non-socketed apical spine. Mesotibia without dorsal pegs. Metacoxa sparsely punctured ventrally, densely so on outer surface of metafemur, with dense and fine setiferous punctures, ventral edge with irregular row of unequal teeth, outer ventral margin with a row of 8-10 teeth, basal tooth not prominent but wider than other teeth; inner basal tooth absent; apical truncation of metatibia forming a curved spine. Tarsi slender.

Metasoma (Fig. 9A, B View Figure 9 ). Petiole not visible dorsally. Gaster lanceolate, longer than mesosoma; Gt1 with weak wrinkles dorsally, setose laterally, the setae progressively longer towards the side (Fig. 9A View Figure 9 ); Gt2-5 smooth, with posterior rows of setiferous punctures, a tuft of sublateral setae longer (Fig. 9A View Figure 9 ); posterior margin of tergites hardly concave; penultimate tergite smooth anteriorly, with moderately coarse setiferous punctures posteriorly; spiracle hardly visible in sublateral position, its aperture much smaller than puncture diameter; syntergum elongate (Fig. 9A, B View Figure 9 ), 1.23 × as long as mesotibia, densely and deeply punctured, with dorsal median ridge (Fig. 9A View Figure 9 ); cerci removed from base of syntergum, situated at half of its length.

Male. Unknown.

Distribution.

Known from Saudi Arabia only in Al-Baha and Asir Regions (Fig. 17 View Figure 17 ).

Host.

Anthaxia (Haplanthaxia) abdita Bílý, 1982 and A. (H.) kneuckeri ssp. zabranskyi Bílý, 1995 ( Coleoptera , Buprestidae ).