Ptomaphagus (s. str.) kuntzeni Sokolowski, 1957

Wang, Cheng-Bin, Ruzicka, Jan, Nishikawa, Masaaki, Perreau, Michel & Hayashi, Yasuhiko, 2016, Revision of the genus Ptomaphagus Hellwig (Coleoptera, Leiodidae, Cholevinae) from Japan, ZooKeys 607, pp. 119-144 : 122-132

Taxon classification Animalia Coleoptera Leiodidae

Ptomaphagus (s. str.) kuntzeni Sokolowski, 1957 Figs 1 A–C; 2 A–F; 3 A–J; 4 A–G

Ptomaphagus (s. str.) kuntzeni Sokolowski 1957: 140 ( Ptomaphagus ; type locality: [JAPAN] Hagi (? Landschaft Jamagutshi, Honshiu); ZMHB); Szymczakowski 1964: 63 ( Ptomaphagus ; female description; taxonomic remarks); Nishikawa 1983: 1 ( Ptomaphagus (Ptomaphagus) ; in check-list); Harusawa and Yamamoto 2000: 242 ( Ptomaphagus ; distribution); Hayashi and Nishikawa 2010: 190 ( Ptomaphagus ; distribution); Perreau 2000: 363 ( Ptomaphagus (s. str.); in catalogue); Perreau 2004: 178 ( Ptomaphagus (Ptomaphagus) ; in catalogue); Nishikawa 2011: 100 ( Ptomaphagus (Ptomaphagus) ; distribution; notes); Nishikawa et al. 2012: 274 ( Ptomaphagus (Ptomaphagus) ; distribution); Perreau 2015: 249 ( Ptomaphagus (Ptomaphagus) ; in catalogue).

Ptomaphagus (s. str.) amamianus Nakane 1963: 42 ( Ptomaphagus ; type locality: [JAPAN] Naze, Amami-Oshima); Hayashi 1969: 2 ( Ptomaphagus ; characteristic figures; distribution); Nishikawa 1983: 1 ( Ptomaphagus (Ptomaphagus) ; in check-list); Perreau 1996: 284 ( Ptomaphagus ; distribution); Perreau 2000: 362 ( Ptomaphagus (s. str.); in catalogue); Perreau 2004: 178 ( Ptomaphagus (Ptomaphagus) ; in catalogue); Hayashi and Nishikawa 2010: 190 ( Ptomaphagus ; distribution); Perreau 2015: 249 ( Ptomaphagus (Ptomaphagus) ; in catalogue). Syn. n.

Material examined.

Type material. Holotype of Ptomaphagus kuntzeni : ♂, [JAPAN] Hagi [ca. 34°24'N, 131°23'E] / Hiller [R. leg., probably collected during 1872-1875 ( Esaki 1935)] // Type // 59051 // Ptomaphagus / kuntzeni Type / det. K. Sokolowski (ZMHB). Holotype of Ptomaphagus amamianus : ♂, HOLOTYPE // [JAPAN] NAZE [ca. 28°22'N, 129°29'E] / AMAMI[- ÔSHIMA] IS. / 4.V.1960 / T. Shibata [leg.] // Ptomaphagus / amamianus Nak. / Det. T. Nakane // NAKANE Coll. / SEHU JAPAN / 1999 // HOLOTYPE / Appended label by / N. Kobayashi / 2008 // 0000005731 / Sys. Ent / Hokkaido Univ. / Japan [SEHU] (HUM). Allotype of Ptomaphagus amamianus : 1♀, ALLOTYPE // [JAPAN] NAZE [ca. 28°22'N, 129°29'E] / AMAMI[- ÔSHIMA] IS. / 4.V.1960 / T. Shibata [leg.] // Ptomaphagus / amamianus Nak. / Det. T. Nakane // NAKANE Coll. / SEHU JAPAN / 1999 // PARATYPE / Appended label by / N. Kobayashi / 2008 // 0000005732 / Sys. Ent / Hokkaido Univ. / Japan [SEHU] (HUM). Paratypes of Ptomaphagus amamianus : 1♂, PARATYPE // [JAPAN] NAZE [ca. 28°22'N, 129°29'E], / AMAMI[- ÔSHIMA] IS. / 4.V.1960 / T. Shibata [leg.] // Ptomaphagus amamianus Nak. / Det. T. Nakane // MHNG / ENTO / 00003333 (MHNG); 1♀, same data as previous except: 00003334 (MHNG); 1♂, PARATYPE // [JAPAN] NAZE [ca. 28°22'N, 129°29'E] / AMAMI[- ÔSHIMA] IS. / 4.V.1960 / T. Shibata [leg.] // Ptomaphagus / amamianus Nak. / Det. T. Nakane // NAKANE Coll. / SEHU JAPAN / 1999 // PARATYPE / Appended label by / N. Kobayashi / 2008 // 0000005733 / Sys. Ent / Hokkaido Univ. / Japan [SEHU] (HUM); 1♀, [JAPAN] PARATYPE // // NAZE [ca. 28°22'N, 129°29'E] / AMAMI[- ÔSHIMA] IS. / 4.V.1960 / T. Shibata [leg.] // Ptomaphagus / amamianus Nak. / Det. T. Nakane // NAKANE Coll. / SEHU JAPAN / 1999 // PARATYPE / Appended label by / N. Kobayashi / 2008 // 0000005734 / Sys. Ent / Hokkaido Univ. / Japan [SEHU] (HUM); 1♂, PARATYPE // [JAPAN] NAZE [ca. 28°22'N, 129°29'E], AMAMI[- ÔSHIMA] IS. / 4.V.1960 / T. Shibata [leg.] // Ptomaphagus amamianus Nak. / Det. T. Nakane // 37-1 [Pl. 37, fig. 1 in Nakane 1963] // NAKANE Coll. / SEMU JAPAN / 1999 // PARATYPE / Appended label by / N. Kobayashi / 2008 // 0000005747 / Sys. Ent Hokkaido Univ. / Japan [SEHU] (HUM).

Additional material.

JAPAN: Honshu: 3♂♂1♀, Botanic Garden [ca. 38°15'N, 140°51'E] / Sendai, Miyagi [Pref.] / 1.IX.1966 / A. NARITA // Ptomaphagus / kuntzeni / Sokolowski / Det. T. Nakane (HUM); 2♂♂, Yoshi-ga-hira [ca. 37°26'N, 139°7'E] / Niigata Pref. / 14.VIII.1987 / leg. M. NISHIKAWA // Trap // alt. 600 m (CMNE); 1♂, ASAMIGAWAKEI / KOKU [ca. 37°13'N, 140°57'E] HIRONOMACHI / FUKUSHIMA [Pref.] 1990.VIII.2 / A. IZUMI leg. (CMNE); 1♀, Mt. TORINOKO-YAMA [ca. 36°41'N, 140°14'E] / 馬頭町 [ Batô-machi], 栃木県 [Tochigi Pref.] / 30.APL.1996 / H. OHKAWA leg. // 14 (CMNE); 1♂, NAGAEDANI [ca. 36°34'N, 136°41'E] / KANAZAWA [Ishikawa Pref.] / 13.V.1948 / S. TAKABA [leg.] (CYHK); 1♂, Mt. TAKAO[-YAMA: ca. 36°27'N 136°45'E] / KAGA [=Ishikawa Pref.] / 2.VII.1961 / Y. Hayashi [leg.] // Ptomaphagus / kuntzeni / Sokolowski? / Det. Y. Hayashi, 1979 (CYHK); 1♂1♀, Tohbu [ca. 36°23'N, 138°21'E], Chiisagata / Nagano [Pref.], JAPAN / 1st, June, 1986 / legit. T. Abe (CMNE); 3♀♀, ISK. [= Ishikawa Pref.] Ishikawa Co. / Shiramine Vil. Shiramine [ca. 36°09'N, 136°37'E] / N. of Mt. Ohnadare / 13. VI– 6.VIII.1988 // (rotten chicken trap) / 630 m, alt. / K. Katsura & / Y. Nishikawa leg. // ♀ (CJRZ); 4♂♂, ISK. [= Ishikawa Pref.] Ishikawa Co. / Shiramine Vil. Katarashi [ca. 36°09'N, 136°38'E] / NW. of Mt. Arigata-yama / 13. VI– 6.VIII.1988 // (rotten chicken trap) / 580 m, alt. / K. Katsura & / Y. Nishikawa leg. // ♂ // Ptomaphagus kuntzeni SOKOLOWSKI / Det. K. Harusawa, 1993 (CJRZ); 1♂1♀, 石川県 [Ishikawa Pref.] 白山市 [Hakusan-shi] / 白山系 [Mts. Hakusan] / 猿壁堰堤 [Sarukabe-entei: ca. 36°06'N, 136°42'E] pit fall traps / 8 月2– 22日2002年 [2-22.VIII.2002] / 保科英人 [Hoshina, Hideto] 採集 [leg.] (CHHF); 1♂, GOZAISHI SPA [ca. 35°43'N, 138°21'E] / YAMANASHI [Pref.] / 12.VIII.1989 / TATEO ITO [leg.] (CYHK); 1♂, 1981 –7– 4 [= 4.VII.1981] / 日川林道上部 [Upper area of Hikawa-rindô: ca. 35°43'N, 138°50'E] // 大ボサツ [Mt. Daibosatsu-rei, Yamanashi Pref.] / 腐敗オサトラップ [decayed bait of carabid trap] // K. HAGA [leg.] (CMNE); 2♂♂, KAMIHIKAWA-RINDO [ca. 35°43'N, 138°50'E] / ([Mt.] DAIBOSATSU[-REI]) / Yamanashi Pref. / Aug. 29-30th 1982 / Y. Shibata leg. (CMNE); 2♂♂, 東京都西多摩郡 [Tokyo, Nishitama-gun] / 檜原村本宿 [Hinohara-mura, Motoshûku: ca. 35°43'N, 139°08'E] // Japan: Honshu / Motoshûku, / Hinohara Mura, / Tokyô To 12.IV.2008 / KAMEZAWA Hiromu leg. // KAMEZAWA Collection (CMNE); 1♂1♀, Mt. Takao[-san: ca. 35°37'N, 139°15'E] / Hachiôji, Tokyo / 16.X.1982 / M. Nishikawa leg. // Trap (CMNE); 4♂♂1♀, Mt. Mitsutoge-yama [ca. 35°32'N, 138°49'E] / 1200 m in alt., trap / Kawaguchiko-machi // Yamanashi Pref., C. / Japan, 19.VIII.1993 / M. Nishikawa leg. (CMNE); 1♂, Dohshigawa [Riv.: ca. 35°32'N, 139°06'E] / Aone, Tsukui Co. / Kanagawa [Pref.], JAPAN / 18th, May, 1986 / T. Abe & A. Sasai [leg.] (CMNE); 2 ♀♀, ( Dôdaira [ca. 35°28'N, 139°10'E]) / E. Tanzawa / Kanagawa Pref. // 11.VII.1993 / leg. M. NISHIKAWA // carrion trap / alt. ca. 1000 m (CMNE); 1♂1♀, same data as previous except: 21.VIII.1993 (CMNE); 1♂, [JAPAN] Honshu, / Kanagawa Pref., Atsugi City, / Shimofurusawa [ca. 35°27'N, 139°19'E], / 23.III.2007. Bait trap. / Takuya FUKUZAWA leg. (CMNE); 1♀, same data as previous except: 22.III.2007 (CMNE); 2♀♀, Idenzawa [ca. 35°26'N, 138°58'E] / Nishitanzawa / Kanagawa [Pref.], Japan // 31.V.-6.VI.2006 / leg. T. Watanabe (CMNE); 1♂, Mt. OHYAMA [ca. 35°25'N, 139°14'E] / Kanagawa [Pref.], Japan / June 15th, 1974 / Coll. Y. Shibata (CMNE); 1♀, Hachikita-kôgen [ca. 35°24'N, 134°32'E] / Muraoka-machi / [A-1] baited-trap / Hyogo Pref. // W Honshu. Japan / 23.XI.2004 / Shigeru Yoshida leg. // 兵庫県村岡町 [Hyogo Pref., Muraoka-machi] / ハチ 北高原 [ Hachikita-kôgen] A-1 / bait-trap / 23.XI.2004 / 吉田 茂 [Yoshida Shigeru] 採集 [leg.] (CMNE); 1♂, Mt. Mikuni-yama [ca. 35°24'N, 138°54'E] / Nishitanzawa / Kanagawa [Pref.], Japan / 3-7.VIII.2003 / T. Watanabe leg. // Flight / Interception / Trap (CMNE); 1♀, (Near OHKURA [ca. 35°24'N, 139°10'E]) / Tanzawa [Mts.], Kanagawa [Pref.] / Apr. 29th, 1973 / Coll. Y. Shibata (CMNE); 3♂♂9♀♀, Kitanisawa 750 m alt. [ca. 35°19'N, 133°33'E] / Mitsukue, Kôfu / Hino-gun, Tottori // Pref., W Japan / 17.VI.2007; b.trap / Y. Fujitani leg. // 鳥取県江府町 [Tottori Pref., Kôfu-chô] / 御机 [Mitsukue] 木谷沢 [Kitanisawa] / bait-trap 750 m (CMNE); 1♂2♀♀, Takahachi [ca. 35°19'N, 138°43'E] on Mt. / Fuji (cola trap) / Shizuoka Pref. // Central Japan / 14.VII.-24.VIII.1996 / M. Nishikawa leg. (CMNE); 1♂, [Mt.] Fudohyama [ca. 35°19'N, 139°11'E] / Nakai-machi / Kanagawa [Pref.], Japan / 3-12.VI.2006 / T. Watanabe leg. (CMNE); 1♂, Hisagi [35°18'N, 139°34'E] / Zushi c. / Kanagawa pref. / 18-23.V.1983. / pit-fall Trap: No.32 / No. 2 (CMNE); 5♂♂12♀♀, Yakôdani [ca. 35°17'N, 134°18'E] / Chizu-machi / Yazu-gun, Tottori / Pref., W Honshu, Jpn / 3.VI.2007 / Y. Fujitani leg. // 鳥取県八頭郡 [Tottori Pref., Yazu-gun] / 智頭町八河谷 [Chizu-machi, Yakôdani] / bait-trap 750 m (CMNE); 6♂♂3♀♀, Near Nontaki waterfall [ca. 35°15'N, 134°08'E] / ca. 800 m in alt. / Aba-son, Tomata-gun / Okayama Pref., Japan / 8.IX.2002; trap / Y. Fujitani leg. (CMNE); 4♂♂2♀♀, Mt. Yamanori-yama [ca. 35°14'N, 133°49'E] / Chûka-son, Maniwa- / ca. 900 m in alt. / gun, Okayama Pref. // W Honshu, W Japan / 11.VII.2004; FIT / Y. Fujitani leg. (CMNE); 2♂♂2♀♀, Daruga-mine [ca. 35°12'N, 134°22'E] / Nishiawakura-son / 1,100 m in alt. / Aida-gun // Okayama Pref. / 7-20.VII.2006; FIT / Akihiko Watanabe leg. (CMNE); 1♂, Akazaigawa Val. [ca. 35°12'N, 134°31'E] / HYOGO Pref. / 12.V.1979 / Y. HAYASHI [leg.] // Ptomaphagus kuntzeni SOKOLOWSKI / Det. Y. HAYASHI, 2014 (CCBW); 1♂1♀, same data as previous except: 15.VII.1979 (MNHA); 1♀, nr. Mt. Suzuga-take [ca. 35°11'N, 136°27'E] / Fujiwara-chô, Mie Pref. / 3.V.2002 / Shiho Arai leg. // Ptomaphagus / kuntzeni / Sokolowski, 1957 / Det. M. Nishikawa, 2014 / #MNIC123909Ch1S (CMNE); 1♀, 千葉県 [Chiba Pref.] 君津市 [Kimitsu-shi] / 郷台畑 [ Gôdaihata] / 猪の川 [Inokawa] / 滝の沢出合 [Takinosawa Deai: 35°11'N, 140°06'E] / malaise traps / 5 月13– 20日1997年 [13-20.V.1997] / 新田 N. [Nitta, N.] 採集 [leg.] (CHHF); 1♂2♀♀, MIZUHO T. [ca. 35°10'N, 135°22'E] / KYOTO, / 22.V.1984 / Y. HAYASHI [leg.] // trap // Ptomaphagus kuntzeni SOKOLOWSKI / Det. Y. HAYASHI, 2014 (1♂ in CCBW and 2♀♀ in CYHK); 1 ♂1♀, same data as previous except: 26.V.1984 (CCBW); 1♂3♀♀, same data as previous except: シズシ [Shizushi: 35°13'N, 135°20'E] // [bar code omitted] (CYHK; MNHA); 1♂, [Mt.] Takiyama [ca. 35°09'N, 134°09'E] / Nagi, Katsuta-gun / Okayama Pref. // W. Honshu, Japan / 9.V.2002 / Y. Fujitani leg. (CMNE); 8♂♂3♀♀, same data as previous except: 7.VII.2002 (CMNE); 3♂♂4♀♀, same data as previous except: 27.X.2002 (CYFO); 1♂, same data as previous except: 10.XI.2002 (CYFO); 1♀, same data as previous except: 23.XI.2002 (CYFO); 1♀, same data as previous except: 29.XII.2002 (CYFO); 19♂♂23♀♀, same data as previous except: 27.VI.2003 (CYFO); 1♀, JAPON KYOTO / Seryô-Tôgé Kyoto [ca. 35°09'N, 135°45'E] / 500-600 m 6.VIII.1980 / Cl. Besuchet [leg.] (MHNG); 1♀, SASAYAMA T. / 雨石山 [Mt. Amaishi-yama: ca. 35°07'N, 135°20'E] / HYOGO [Pref.], / 28.IV.1984 / Y. HAYASHI [leg.] (CYHK); 2♂♂3♀♀, nr. Rashomon-daiichi- / do Cave [ca. 34°56'N, 133°33'E], Niimi-shi. / 370 m in alt. / Okayama Pref. // W Japan (carrion trap) / 18.VIII.2001 / Y. Fujitani leg. (CMNE); 1♂1♀, Shiramizu [ca. 34°58'N, 134°15'E] / Mimasaka-shi / [carrion baited-trap] / Okayama Pref. // W Honshu, Japan / 7.V.2005 / Y. Fujitani leg. (CMNE); 4♂♂1♀, Mumyôdani [ca. 34°56'N, 133°27'E], ca. 350 m / Tetta-chô, Atetsu- / gun, Okayama Pref. // W Honshu, W. Japan / 25.IV.2004; b. trap / Y. Fujitani leg. (CMNE); 1♀, 箕面市下止々呂美 [ Minô-shi, Shimotodoromi: ca. 34°52'N, 135°27'E] / (OSAKA pref.) / 4.VIII.1992 / leg. 齋藤琢己 [Saito Takumi] // Ptomaphagus / kuntzeni / SOKOLOWSKY / Det. Y. HAYASHI, 1993 (CYHK); 1♂4♀♀, Kanahira National / Forest, [ca. 34°47'N, 133°37'E] Takahashi- / [carrion baited-trap] / shi, Okayama Pref. // W Honshu, Japan / 16.IV.2006 / Y. Fujitani leg. (CMNE); 1♂, Anatoyama-jinja [ca. 34°39'N, 133°29'E] / 450 m alt., Kawakami / Okayama Pref. // Honshu, W Japan / 22.IV.2001; trap / Y. Fujitani leg. (CMNE); 1♀, same data as previous except: 8.IV.2001 (CMNE); 1♀, same data as previous except: 16.IV.2001 (CMNE); 1♂3♀♀, same data as previous except: 28.IV.2001 (CMNE); 1♀, Ômukai-rindô [ca. 34°29'N, 132°07'E] / Chôjabara, Yoshiwa / 880 m in alt. / Hatsukaichi-shi // Hiroshima Pref., W / Honshu, Jpn. 28.IV. / 2007, M. Tagami leg. // 広島県廿日市市 [Hiroshima Pref., Hatsukaichi-shi] / 吉和長者原大向林道 [Yoshiwa, Chôjabara, Ômukai-rindô] / FIT 880 m / 28.IV.2007 / 田上雅生 [Tagami Masao] 採集 [leg.] (CMNE); 1♂1♀, OSAKA Pr. Tondabayashi c. / Kannobi Kongô Colony [ca. 34°29'N, 135°35'E] / 12-26.IV.1994 / leg. K. Harusawa // pit fall trap: / rotten squid // Ptomaphagus kuntzeni Sokolowski (CMPR); 6♂♂3♀♀, Kasayama, ca. 50 m / Hagi-shi, / ca. 34°26'N, 131°24'E / Yamaguchi Pref. // W Honshu, W Japan / 29.VI.2002; trap / Y. Fujitani leg. (CMNE); 2♀♀, same data as previous except: 5.VII.2002 (CMNE); 2♂♂2♀♀, Mt. Makio[-san: ca. 34°23'N, 135°30'E] / OSAKA [Pref.] / 4.VIII.1983 / F. KIMURA [leg.] (CYHK); 1♂, [OSAKA Pr.] / Kawachinagano / 天見 [Amami], alt. 260 m / 6-14.X.1981 / K. Harusawa / Y. Nishikawa leg. // 出合 [Deai]~流谷 [Nagaredani] / 流谷八幡境内 [Nagaredani-hachiman shrine’s area: ca. 34°23'N, 135°35'E] / 腐肉 [carrion] Trap (牛 [beef]) // Ptomaphagus / kuntzeni / SOKOLOWSKI / Det. Y. HAYASHI, 1990 (CYHK); 1♂, Kamisuga [ca. 34°23'N, 136°22'E] / Ôdai-machi, Taki- / gun, Mie Pref. // Honshu, Japan / 22-9.V.2005 / Katsumi Akita leg. (CMNE); 3♂♂2♀♀, Kanoashikôchi [ca. 34°22'N, 131°57'E] / Muikaichi-machi / 450 m in alt. / Shimane Pref. // W Honshu, Japan / 10.VI.2005 / Y. Fujitani leg. // 六日市町 [Muikaichi-machi] / 鹿足河内 [ Kanoashikôchi] (CMNE); 1♂, HASE [ca. 34°19'N, 135°48'E] / YAMATO [=Nara Pref.] / 5.VI.1966 / Y. HAYASHI [leg.] // Ptomaphagus / kuntzeni (CYHK); 1♂4♀♀, Nodani [ca. 34°17'N, 131°38'E], Tokuji-chô / Saba-gun / Yamaguchi Pref. // W Honshu, W Japan / 21.IV.2004; b. trap / Y. Fujitani leg. (CMNE); 2♀♀, mouth of Nakao-dô / Cave [ca. 34°16'N, 131°17'E]. Aokage / Shuhô, Yamaguchi // Pref., W Japan / 28.VII.-4.VIII.2002 / Y. Fujitani leg. (CMNE); 3♂♂5♀♀, Akiyoshidai [ca. 34°16'N, 131°18'E] / Shuho-machi / Yamaguchi Pref. // W Honshu, W Japan / 12.IV.2002; trap / Y. Fujitani leg. (CMNE); 1♂2♀♀, Mt. Nagano-yama [ca. 34°16'N, 131°52'E] / 1,010 m in alt. / b.trap / Kano-machi // Yamaguchi Pref. / Japan, 4.VI.2005 / Y. Fujitani leg. (CMNE); Japan. / G. Lewis. / B. M. 1926-369. // Chiuzenji [ca. 36°46'N, 139°28'E]. // Ptomaphagus / sibiricus Jean. / Jeannel det. // Ptomaphagus / kuntzeni ♀ / det. WANG C.-B., 2016 (BMNH). Kyushu: 1♂, Hata [ca. 33°48'N, 130°46'E] / Yahata-City [Fukuoka Pref.] / 3.V.1965 / coll. M. Ueda (HUM); 1♂, 福岡県 [Fukuoka Pref.] 添田町 [Soeda-cho] / 英彦山 [Mt. Hiko-san: ca. 33°29'N, 130°54'E] / 5月2日1983年 [2.V.1983] / 野村 S. [Nomura S.] 採集 [leg.] (CHHF); 1♀, (Kinsenji [ca. 32°58'N, 130°05'E], Mt. Tara[-dake]) / Nagasaki Pref. / Kyushu, JAPAN / 10.V.1983 / S. Imasaka leg. // Ptomaphagus (s. str.) / kuntzeni Sokolowski, 1957 / Det. M. Nishikawa, 2010 / ♀ (CMNE); 1♂, Mt. Gokabaru-dake [ca. 32°57'N, 130°04'E] / (1,058 m in alt.) / Nagasaki Pref. // NW Kyushu, Japan / 11-12.V.1991 / M. Nishikawa leg. // carrion trap / alt. ca. m (CMNE); 2♂♂, Unzen-bessho [ca. 32°44'N, 130°15'E] / Nagasaki Pref. / 1.VI.1983 / leg. S. Imasaka (CMNE); 1♂, Hagi [ca. 32°32'N, 130°56'E], Gokanoshô / Kumamoto Pref. / 18.VI.1984 / leg. S. Imasaka (CMNE); 1♀, Takao [ca. 32°28'N, 131°08'E], Shimofukura / Shiiba, Miyazaki Pref. / Kyushu, SW Japan / 26.VII.2015 / Takashi Watanabe leg. // Ptomaphagus ( Ptomaphagus ) / kuntzeni Sokolowski, 1957 / Det. M. Nishikawa, 2016 / MNIC124950Ch1S♀ (CMNE); 1♂, Inao dake [ca. 31°07'N, 130°53'E] / Tashiro cho / Kagoshima Pref. / 14.VII.1985 / T. TANABE leg. // alt. 300 m // Trap // Ptomaphagus / kuntzeni / Sokolowski / Det. T. Nakane (HUM); 1♂, same data as previous except: 575 m (HUM); 1♀, same data as previous except: 610 m (HUM). Ryukyus: Amami-Ôshima Is.: 1♀, 鹿児島県 [Kagoshima Pref.] / 奄美市 [Amami-shi] / 龍郷町 [ Tatsugô-chô] / 芦徳 [Ashitoku: ca. 28°25'N, 129°36'E] / 05.V.2011 / 稲垣政志 [Inagaki Masashi] Leg. (CYHK); 1♂1♀, (JAPAN) Kagoshima pref., / Amami-Ohshima Is., / Honcha pass. [ca. 28°23'N, 129°33'E] / 11-14.IV.2007 / T. FUKUZAWA et al. [leg.] (CMNE); 1♀, NAZE [ca. 28°22'N, 129°29'E] / AMAMI[- ÔSHIMA] IS. / 4.V.1960 / T. Shibata [leg.] // Ptomaphagus / ohshimensis / NAKANE / Det. Y. HAYASHI, 1969 (CYHK); 2♀♀, same data as previous but no det. label (CYHK); 1♀, same data as previous except: // Ptomaphagus / amamianus / NAKANE / Det. Y. HAYASHI, 2009 // Collection of / Y. HAYASHI // [bar code omitted] (MNHA); 2♂♂4♀♀, Ôganeku / 28.3598N 129.3403E [ca. 28°21'N, 129°20'E]; 260 m alt. / Yamato-son / Amami-Ôshima Is., Ryukyus / Kagoshima Pref., SW Japan / 27.II.-2.III.2016; trap / M. Nishikawa leg. (CMNE); 4♀♀, nr. Hôkoku-jinja, ca. 350 m / 28.362331N 129.482009E [ca. 28°21'N, 129°28'E] / Amami-shi / Amami-Ôshima Is. / Kagoshima Pref., Ryukyus / SW Japan / 27.II.-2.III.2016; trap / M. Nishikawa leg. (CMNE); 3♂♂4♀♀, around Ôkawa Dam, ca. 28°20'N, 129°29'E, 130 m alt., Amami-shi, Amami-Ôshima Is. / Kagoshima Pref., Ryukyus / SW Japan / 27.II.-2.III.2016; trap / M. Nishikawa leg. (CMNE); 1♂, Toen, nr. Amami ForestPolis / 28.3176°N 129.3297°E [ca. 28°19'N, 129°19'E]; 180 m alt. / Yamato-son, Amami-Ôshima Is. / Ôshima-gun, Kagoshima Pref. / Ryukyus, SW Japan / 3-10.III.2015; bait trap / M. Nishikawa leg. (CMNE); 11♂♂18♀♀, (JAPAN) Kagoshima Pref., / Amami-Ôshima Is., / Yamato vil., Ôdana. 310 m alt. / N28°18'40.2 E128°55'32.2 / 5.III.2008. Carrion Baited trap. / Takuya FUKUZAWA leg. (CMNE); 8♂♂5♀♀, Naon-kengyôzôrin Forest [ca. 28°18'N, 129°20'E] / 名音県行造林 [ Naon-kengyôzôrin Forest]; alt. 349 m / 28.3058°N, 129.3371°E / Yamato, Amami-Ôshima Is. // Ôshima-gun, Kagoshima Pr. / Ryukyus, SW Japan / 2-10.III.2015; bait trap / M. Nishikawa leg. // Ptomaphagus ( Ptomaphagus ) / amamianus Nakane, 1963 / Det. M. Nishikawa, 2015 (CMNE); 1♂, Mt. Yuwan[-dake: ca. 28°17'N, 129°19'E] / Amami-Ohshima Is / 29.III.1999 // Ptomaphagus amamianus NAKANE / Det. Y. HAYASHI, 2014 (CCBW); 1♀, same data as previous except: 28.IV.2000 / 江本健一 [Emoto, Ken’ichi] 採集 [leg.] // ♀ (CMNE); 1♀, same data as previous except: 7-12.V.2006 / T. Watanabe leg. (CMNE); 1♂3♀♀, same data as previous except: // 650-680 m, / 18-26.III.2010, baited / pitfall traps, forest, / Tomáš Lackner leg. (CJRZ); 9♂♂9♀♀, (JAPAN) Kagoshima Pref., / Amami-Ôshima Is., sumiyou Vil., santarou Touge Pass, 318 m alt. / N28°17'03.9 E129°25'16.2 / Carrion Baited Trap. 6.III.2008. / Takuya FUKUZAWA leg. (CMNE); 2♂♂1♀, same data as previous except: 12-14.IV.2007 (CMNE); 1♂1♀, same data as previous except: 13-16.IV.2007. FIT / T. FUKUZAWA et al. [leg.] (CMNE); 4♂♂2♀♀, 鹿児島県大島郡 [Kagoshima Pref., Ôshima-gun] / 宇検村 [Uken-son] ヤクガチョボシ 山麓 [foot of Mt. Yakugachoboshi-yama: ca. 28°15'N, 129°21'E] // Japan; Ryukyu / foot of Yakugachoboshi-yama, / Uken Son, Oshima Gun, / Kagoshima Ken // (Is. Amami-o-shima) / 27.II.2004 / KAMEZAWA, Hiromu leg. (CMNE); 1♂, HATSUNO [ca. 28°15'N, 129°22'E] / Amami[- Ôshima] Is. / 31.III.1969 / H. NOMURA [leg.] // Ptomaphagus amamianus NAKANE / Det. Y. HAYASHI, 2014 (CCBW); 1♂, same data as previous except: 1.IV.1967 (CYHK); 1♀, same data as previous except: 3.IV.1967 (CYHK); 1♂, same data as previous except: 4.IV.1967 (CYHK); 2♂♂1♀, same data as previous except: 3.IV.1969 // K. TANIZAWA [leg.] (CCBW); 2♂♂1♀, same data as previous except: // Ptomaphagus / amamianus NAKANE / Det. Y. HAYASHI, 1993 (CMNE); 1♂1♀, same data as previous except: 31.III.1969 / H. NOMURA [leg.] (CYHK); 4♂♂, same data as previous except: 1.IV.1969 (CYHK); 4♂♂1♀, same data as previous except: 2.IV.1969 (CYHK); 1♂1♀, same data as previous except: 3.IV.1969 (CYHK); 7♂♂3♀♀, same data as previous except: K. Tanizawa [leg.] (CYHK); 1♀, same data as previous except: 5.V.1969 / Y. MAEDA [leg.] (CMNE); 1♀, Nishinakama [ca. 28°15'N, 129°24'E] / AMAMI[- ÔSHIMA] Isl. / 5.IV.1969 / K. TANIZAWA [leg.] (CYHK); 3♂♂3♀♀, nr. Yakugachi Tunnel / 28.2295N 129.3599E [ca. 28°13'N, 129°21'E]; 30 m alt. / Kamiyakugachi, Amami-shi / Amami-Ôshima Is., Ryukyus / Kagoshima Pref., SW Japan / 27.II.-2.III.2016; trap / M. Nishikawa leg. (CMNE); 1♀, (由井岳 [Mt. Yui-dake: ca. 28°11'N, 129°18'E]) / 瀬戸内 [ Setouchi-chô] 奄美大島 [ Amami-Ôshima Is.] / 25.IV.2000 / 江 本健一 [Emoto Ken’ichi] 採集 [leg.] // ♀ (CMNE); 6♂♂, Aminoko-tôge Pass / 28.1893°N 129.3659°E [ca. 28°11'N, 129°21'E]; 350 m alt. / Setsuko, Setouchi-chô / Amami-Ôshima Is. / Ôshima-gun, Kagoshima Pref. / Ryukyus, SW Japan / 6-10.III.2015; bait trap / M. Nishikawa leg. (CMNE); 1♂3♀♀, 鹿児島県 [Kagoshima Pref.] / 大島郡 [ Ôshima-gun] / 瀬戸内町 [ Setouchi-chô: ca. 28°08'N, 129°18'E] / 05.V.2011 / 稲垣政志 [Inagaki, Masashi] Leg. (CYHK); 1♂, IKARI [unlocated] / AMAMI[- ÔSHIMA] IS. / 18.V.1960 / T. Shibata [leg.] (CYHK). Ryukyus: Kume-jima Is.: 3♀♀, Shirase-gawa [Riv.: ca. 26°20'N, 126°46'E] / Gushikawa-son / Kume-jima Is. // Ryukyus, SW Japan / 15-17.III.1998 / M. Maruyama leg. (CMNE). Ryukyus: Okinawa-hontô Is.: 1♂1♀, [Okinawa: JAPAN] / Uka-rindô [ca. 26°48'N, 128°14'E], alt. 250- / 300 m, Kunigami vill. / 8.II.2009 / Takashi Kurihara leg. // ベイトトラップ [bait trap] / 鳥の手羽元 [fowl wing sticks] // Ptomaphagus / kuntzeni ? / Y. HAYASHI, 19 2014 (CYHK); 2♂♂, same data as previous except 7.II.2009 (CYHK); 2♂♂, Yona [ca. 26°45'N, 128°12'E] (Ohkuni 5) / Okinawa-jima Is. / Ryukyus // SW Japan / 25-27.IV.1996 / M. Nagano leg. (CMNE); 1♀, Nishime-dake [ca. 26°48'N, 128°16'E]. OKN [= Okinawa Pref.] / Date: 14.X.2002 / K. MASUMOTO leg. (CMNE); 2♀♀, [Okinawa: JAPAN] / Hama-rindô [ca. 26°43'N, 128°09'E], alt. 50- / 100 m, Kunigami vill. / 8.II.2009 / Takashi Kurihara leg. // ベイトトラップ [bait trap] / 鳥の手羽元 [fowl wing sticks] (CYHK); 4♂♂, (JAPAN, Ryukyus) / Okinawa pref. / Okinawa Is. / Kunigami Vil., Hiji [ca. 26°43'N, 128°10'E] / 13.III.2009. / Takuya FUKUZAWA leg. (CMNE); 1♂, 与那覇岳 [Mt. Yonaha-dake: ca. 26°42'N, 128°13'E] / 沖縄本島 [ Okinawa-hontô Is.] / 14.iv.2000 / 保科英人 [Hoshina, Hideto leg.] / 腐肉トラップ [carrion trap] // 2 (CMNE); 7♂♂5♀♀, bank of Haneji-ô-kawa / Riv. [ca. 26°36'N, 128°01'E], 20-40 m in alt. / Nago-shi, Okinawa- // jima Is., Ryukyus / (c. trap) 18.IV.1993 / R. Yakita leg. (CMNE); 13♂♂7♀♀, same data as previous except: 8-18.IV.1993 (CMNE); 14♂♂9♀♀, (JAPAN, Ryukyus) / Okinawa Pref. / Okinawa Is. / Nago City, Genka. / 26°36'N, 128°04'E // 10-13.III.2009. / by flight Intercept Trap / T. Fukuzawa, T. Ishikawa & M. Kishi leg. (CMNE). Ryukyus: Tokuno-shima Is.: 1♂, (JAPAN) Kagoshima Pref. / Tokuno-shima Is., / Amagi Town, / Mt. Yamatogusuku-san [ca. 27°48'N, 128°55'E] / 29.II.-4.III.2008. (FIT) Takuya FUKUZAWA leg. (CMNE); 85♂♂67♀♀, (JAPAN) Kagoshima Pref. / Tokuno-shima Is., / Amagi Town, / Mt. Sankyô-dake, 185 m alt. / N27°46'13.6, E128°57'18.0 / Carrion Baited Trap. 4.III.2008. / Takuya FUKUZAWA leg. (CMNE); 4♂♂3♀♀, same data as previous except: 27.II.-4.III.2008. (FIT) (CMNE). Sadoga-shima Is.: 1♂, Mt. Donden-yama [ca. 38°07'N, 138°22'E] / Sado Is., Niigata / Pref., Honshu, Japan // 30.IV.-26.V. / 1990 / leg. M. NISHIKAWA // carrion trap / alt. ca. 200 m (CMNE). Shikoku: 1♀, [SHIKOKU] / Komenono [ca. 33°58'N, 132°51'E] / Matsuyama [Ehime Pref.] / 25.IV.1993 / Lizhen Li leg. (EUM); 1♂, (Ehime: Japan) / Aonamimachi [ca. 33°53'N, 132°51'E] / Matsuyama-shi / 24.V.2006 / Shôma Sejima leg. (EUM); 3♀♀, same data as previous except: 2.VI.2006 (EUM); 2♀♀, Minokoshi on Mt. / [ca. 33°51'N, 134°05'E] / Tsurugi-san / ca. 1400 m in alt. / Tokushima Pref. // Shikoku, Japan / 25.VII.2004; trap / Y. Fujitani leg. // 德島県東祖谷山村 [Tokushima Pref., Higashiiyayama-son] / 見の越 [Minokoshi] / bait- trap (CMNE); 2♂♂2♀♀, 瓶が森 [Mt. Kamegamori: ca. 33°47'N, 133°11'E] (alt. 1670 m) / 高知県吾川郡いの町 [ Kôchi Pref., Agawa-gun, Ino-chô] / 寺川 [Terakawa] 10.V.-8.VIII.2009 / 吉田 正隆 [Yoshida Masataka] 採集 [leg.] // Mt. Kamegamori (alt. 1670 / m) Terakawa Ino-chô / Agawa-gun Kôchi-Pref. / 10.V.-8.VIII.2009 / Masataka YOSHIDA leg. // 鳥ガラトラップ [fowl bone trap] / 地中 [underground] (50 cm) (CMNE); 2♂♂1♀, [SHIKOKU] / Mt. Ishizuchi [ca. 33°44'N, 133°06'E] / Ehime Pref. / 27.VIII.1990 / alt. 800 m / S. Takano leg. (EUM); 2♂♂, same data as previous except: 13.IX.1990 / alt. 500 m (EUM); 1♂, EHIME [Pref.]; Japan / Mt. Saragamine [ca. 33°43'N, 132°53'E] / Shigenobu-Town / 22.IV.1999 / Tatsuya Kan leg. (EUM); 4♀♀, 156 JAPAN, Shikoku, / Jshizuchi [Ishizuchi] Mt. Nat. Park, / OMOGO Valley, [ca. 33°42'N, 133°05'E] 700 m, / S. & J. Peck leg. // 158 mega carrion traps, / mixed warm temperate / forest, 18-25.viii.1980 (CJRZ); 1♀, Nishidani [ca. 33°31'N, 132°57'E] / Yanadani, Ehime [Pref.] / 6-8.V.1994 / Sakai, Li, Aita leg. / (bait-trap) // No. 8 (EUM); 4♂♂2♀♀, (SHIKOKU) / Odamiyama [ca. 33°31'N, 132°53'E] / Ehime Pref. / 22.VIII.1984 / E. Yamamoto [leg.] (EUM); 1♂3♀♀, same data as previous except: 14.VIII.1984 / 山本栄治 [Yamamoto Eiji] 採集 [leg.] (EUM); 1♂2♀♀, Komi [ca. 33°30'N, 132°57'E] / Yanadani, Ehime [Pref.] / 6-8.V.1994 / Ohbayashi, Nishino, Okada [leg.] / (by bait-trap) // No. 2 (EUM); 1♂1♀, same data as previous except: No.1 (EUM); 4♂♂5♀♀, Tengu-Kôgen [ca. 33°28'N, 133°00'E] (alt. 1280 m) / Kumakôgen-chô Kamiuke- / na-gun Ehime-Pref. / 9.V.-25.VII.2010 (Fowl trap) / Masataka YOSHIDA leg. // 愛媛県上浮穴郡 [Ehime Pref., Kamiukena-gun] / 久万高原町天狗高原 [ Kumakôgen-chô, Tengu-Kôgen] / (alt. 1280 m) 9.V.-25.VII. / 2010 (鶏ガラトラップ [chicken bone trap]) / 吉田正隆 [Yoshida Masataka] 採集 [leg.] (CYHK); 1♀, Nagano [ca. 33°27'N, 132°56'E] / Yusuhara-chô [ Kôchi Pref.] / 1-2.X.1994 / Y. Utsunomiya [leg.] / (by bait-trap) // No. 9 (EUM). Shimokoshiki-jima Is.: 1♀, Mt. Odake [ca. 31°43'N, 129°44'E] / Is. Shimokoshiki[-jima] / Kagoshima-pref. / 20.VI.1982 / S. Imasaka leg. (CMNE).

Redescription.

Male. EBL: 3.5-4.4 mm (3.5 mm in holotype of Ptomaphagus kuntzeni ). Length of different body parts: HL: AL: PL: ELL = 0.6: 1.2: 1.0: 2.2 mm; width: HW: EW: PW: ELW = 1.0: 0.1: 1.6: 1.6 mm. Proportion of antennomeres from base to tip in µm (length × width): 167 × 84, 118 × 68, 114 × 74, 67 × 80, 76 × 99, 53 × 114, 96 × 126, 42 × 128, 92 × 142, 97 × 138, 197 × 115.

Habitus (Fig. 1A, B) elongated oval, regularly convex and sublustrous. Well pigmented: mostly brown to dark brown, head darker; mouthparts, basal three or four antennomeres and apical half of ultimate antennomere, protarsi, and apical parts of meso- and metatarsi more or less paler. Dorsum continuously clothed with fine, recumbent, yellowish pubescence. Insertions of pubescence on dorsal surfaces of pronotum, elytra and femora aligned along transverse striolations; interspace between two striolations glabrous.

Head quite transverse, HW/HL = 1.6. Clypeofrontal suture absent. Clypeus with anterior margin slightly rounded. Compound eyes well developed, EW/HW = 0.1. Antennae (Fig. 3A) slender, AL/HW = 1.2; antennomere III as long as II; VI with length/width = 0.5; XI longest, elongated pear-shaped.

Pronotum (Fig. 3B) transverse, widest just before hind angles, PW/PL = 1.5. Sides gently arched, gradually narrowing from posterior to anterior; hind angles drawn out, acute and sharp. Posterior margin widely protruding in the middle part, distinctly emarginate near hind angles.

Elytra oval, widest near basal 2/7, ELL/EW = 1.4. Sides weakly arched, gradually narrowing from widest part to apices, which widely rounded (Fig. 3G). Sutural striae present. Metathoracic wings fully developed.

Prolegs robust, with basal three protarsomeres (Fig. 3C) moderately expanded: TW/BTW = 1.4. Protibiae (Fig. 3E) distinctly expanded towards apex. Profemora (Fig. 3E) broad. Mesotibiae arcuate, mesotarsi simply linear. Metatibiae slender and straight.

Abdominal ventrite VIII (Fig. 3I) round at posterior edge and with an inconspicuous median notch. Spiculum gastrale of genital segment (Figs 2C, F; 3J) with approx. 1/3 of length protruding beyond anterior edge of epipleurite IX.

Aedeagus stout and wide, with median lobe gradually narrowing towards an oblong apex and terminated by an obtusely rounded knob in dorsal view (Figs 2A, D; 4A); opening of genital orifice situated on dorsal surface, deeply cut inwards on left edge of median lobe at subapex. Ventral surface of the apex of the median lobe (Fig. 4B, D) inserted with 5 ventrally oriented setae on the left side and 4 ventrally oriented setae on the right side; parameres narrow, reaching almost to apical 1/5 of median lobe, each apex (Fig. 4E) with 2 lateral setae and 1 apical seta relatively shorter. In lateral view (Fig. 2B, E), median lobe distinctly thick, regularly bent ventrad and gradually tapering towards a subround apex. Endophallus with stylus quite slender, a cheliform complex below the base of stylus, and a circular complex at the basal region.

Female. Similar to male in general appearance (Fig. 1C), including elytral apices (Fig. 3H), but distinguished by the following characteristics: protarsi (Fig. 3D) simply linear; protibiae (Fig. 3F) narrower at apex; ventrite VIII (Fig. 4F) almost narrowly rounded at posterior edge; genital segment as shown in Fig. 4G; spermatheca (Fig. 4G) C-shaped in distal part, not coiled in proximal part.

Distribution.

China (Taiwan), Japan (Fig. 8),?Myanmar.

Remarks.

Varying body size is not an unexpected intraspecific variation in Ptomaphagus species, and occurs in common European species such as Ptomaphagus (s. str.) sericatus (Chaudoir, 1845) and Ptomaphagus (s. str.) varicornis (Rosenhauer, 1847). Although the holotype and paratypes of Ptomaphagus (s. str.) amamianus (Fig. 1B, C) are larger than the holotype of Ptomaphagus (s. str.) kuntzeni (Fig. 1A), this does not prevent us from synonymising the two species because of their identically shaped aedeagus (Fig. 2A, B, D, E).

Ptomaphagus (s. str.) kuntzeni somewhat resembles Ptomaphagus (s. str.) sibiricus in general appearance, but the former has antennomere III as long as II (Fig. 3A), antennomere VI with length/width = 0.5 and elytral apices (Fig. 3G, H) widely rounded; while the latter has antennomere III a little shorter than II (Fig. 7A), antennomere VI with length/width = 0.4 and elytral apices (Fig. 7B) narrowly rounded.

It should be noticed that Szymczakowski (1964) described a single female specimen from Kambaiti, Myanmar (coll. Naturhistoriska Riksmuseet, Stockholm) as Ptomaphagus (s. str.) kuntzeni . We concur with the opinion of Nishikawa (2011) that the occurrence of Ptomaphagus (s. str.) kuntzeni in Myanmar is dubious because of the wide geographical gap and the discrepancies of the morphological description of Szymczakowski (1964) with specimens from Japan.

According to the present data, this species is one of the most widespread cholevines in Japan, known from Honshu, Shikoku, Kyushu and Ryukyus. However, we have not yet examined any specimens from the northernmost area of Honshu (above the 38th parallel) or from the southern Kume-jima Island in Ryukyus (Fig. 8). The species is recorded herein from Sadoga-shima Island, Shimokoshiki-jima Island and Kume-jima Island for the first time. Incidentally, no Ptomaphagus species have been recorded from Hokkaido to date. Moreover, Perreau (1996) reported the species from Taiwan Island under the name Ptomaphagus (s. str.) amamianus . We will deal with this area in a next paper devoted to Ptomaphagus from Taiwan.

Collecting methods for the material examined mostly indicate a necrophagous association, such as traps baited with decaying animal matter.