Thysananthus gottschei var. continuus, Sukkharak. A. Portion, 2015

Sukkharak, Phiangphak, 2015, A systematic monograph of the genus Thysananthus (Lejeuneaceae, Marchantiophyta), Phytotaxa 193 (1), pp. 448-450 : 448-450

8b. Thysananthus gottschei var. continuus , var. nov. Type: MALAYSIA. Sarawak: “Lunda”, without date, Micholitz s.n. (holotype G!). The name of the new variety refers to the free margin of lobules continuing into the lobe ventral margin over almost the entire length of the ventral margin, and sometimes into the dorsal margin of the lobes .

Keel with appendages on one side where leaves and underleaves are free and opposite to adnate ones, always curved towards the stems. Lobule free margin continuing into both ventral and dorsal lobe margins. Underleaves apex strongly recurved. Fig. 18 .

Distribution and ecology:— Malaysia and Philippines; without ecological information.

Representative specimens:— Philippines. ILOILO: Panay, Robinson BS18185 (L).

Taxonomic notes:— The distinguishing characters of this variety are (1) presence of appendage on the keel, on the side where leaves and underleaves are free and opposite to the adnate ones (always curved towards the stems), (2) free margin of lobule continuing into the lobe ventral margin over almost the entire length of the ventral margin, and sometimes into the dorsal margin of the lobes, and (3) strongly recurved underleaf apex.

Figures

FIGURE 18. Thysananthus gottschei var. continuus Sukkharak. A. Portion of shoot, ventral view. B. Leaves, ventral view: a = appendage. C. Underleaves, ventral view. D. Female bracts, ventral view: a = appendage. E. Female bracteole, ventral view. F. Perianth, dorsal view. G. Perianth, ventral view. H. Margin cells of leaf lobe. I–J. Leaf lobule apices. K. Leaf lobule base. (All from the lectotype).