Lagenosoma geminatum, Winterton, 2023

Winterton, Shaun L., 2023, Wasp-mimicking soldier flies of the Australian Region: revision of Ampsalis Walker, Elissoma White and Lagenosoma Brauer (Stratiomyidae: Clitellariinae), Zootaxa 5246 (1), pp. 1-63 : 38

Lagenosoma geminatum sp. n.

( Figs 16E–H ; 20 ; 27C, D ; 30B ; 37 )

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Diagnosis. Antennae relatively short, flagellum black, flagellomeres VI and VII each distinctly longer than lengths of flagellomeres IV and V; flagellomere VII ca. 1/3 length of flagellomere VIII; male head relatively small; male postocular ridge broadly carinate; scutum and scutellum orange with black punctate setal bases; pterostigma yellow; hind leg longer than other legs.

Description. Body length = 8.0 mm. Head. Yellow to orange, frons and face rounded with short, pale setal pile, female sometimes with paired triangular black marks above antenna; parafacial with silver pubescence along eye margin; ocellar tubercle black; postocular ridge yellow, strongly carinate in male; occiput flat, yellow, covered with short, black, setal pile; antenna short (length = 1.9 mm), scape, pedicel and flagellomeres I–III dark yellow with dark microtrichia, flagellomeres VI–VIII black and shallow plumose, flagellomeres VI and VII each distinctly longer than lengths of flagellomeres IV and V, ratio of length of flagellomere VII to VIII is 1: 3.3, flagellomere VIII narrow, shallowly plumose at most. Thorax. Dark yellow to orange with postpronotum, scutum and scutellum with black punctate setal bases; coxae and legs yellow, tarsi suffused brown to black, hind tarsi darker; legs elongate, hind legs noticeably longer; haltere yellow; wing hyaline, venation and pterostigma yellow to brown. Abdomen. Slightly narrowed basally in both sexes, ovoid shaped and not petiolate, segment 4 widest and equal to width of scutum; segment 1 with lateral flange well developed; abdomen yellow with short black setal pile. Male genitalia. Not dissected, but externally epandrium quadrangular; gonocoxites relatively elongate and posterolateral processes broad; gonostylus relatively small. Female genitalia. Not dissected.

Etymology. The species epithet is derived from the Latin geminatum —double, pair, referring to the similarity with L. eklemmum sp. n.

Comments. Lagenosoma geminatum sp. n. is closely related to L. eklemmum sp. n. and females are similar in appearance; major differences are in wing pterostigma colour, size of the male head and width of postocular ridge, and leg length in both sexes. The scutum is differently shaped in profile in both sexes as well. This species is known from coastal central Queensland.

Type material— Holotype male, AUSTRALIA: Queensland: Gayndah [-25.6108, 151.5969], Masters ( QM).

Paratypes. AUSTRALIA: Queensland: 1 female, Boomer Range, Python Scrub, site 5 [-23.1993, 149.7343], 28–29.X. 1999, 240 m Monteith , Cook , Burwell , Evans , vine scrub ( QM) ; 1 female, Lake Barrine [-17.2448, 145.6393], 18.XII.1959, L.J. Harris ( QM) .

Figures

FIGURE 16. A. Lagenosoma eklemmum sp. n., male, lateral view; B. Lagenosoma eklemmum sp. n., male, oblique view; C. Lagenosoma eklemmum sp. n., female, oblique view; D. Lagenosoma eklemmum sp. n., female, lateral view; E. Lagenosoma geminatum sp. n., male, lateral view; F. Lagenosoma geminatum sp. n., male, anterolateral view; G. Lagenosoma geminatum sp. n., female, oblique view; H. Lagenosoma geminatum sp. n., female, lateral view.

FIGURE 20. Stylised illustrations of the antenna of Eudmeta, Ampsalis, Elissoma and Lagenosoma.

FIGURE 27. Heads of Elissoma and Lagenosoma. A. Lagenosoma eklemmum sp. n., male; B. Lagenosoma eklemmum sp. n., female; C. Lagenosoma geminatum sp. n., male; D. Lagenosoma geminatum sp. n., female; E. Lagenosoma pictum Brauer, male; F. Lagenosoma pictum Brauer, female; G. Lagenosoma propinquum Brauer, male; H. Lagenosoma propinquum Brauer, female; I. Lagenosoma varigatum sp. n., male; J. Lagenosoma varigatum sp. n., female; K. Elissoma hauseri sp. n., male; L. Lagenosoma propius sp. n., male.

FIGURE 30. Lagenosoma scutellums. A. Lagenosoma eklemmum sp. n., female; B. Lagenosoma geminatum sp. n., male; C. Lagenosoma pictum Brauer, male; D. Lagenosoma pictum Brauer, female; E. Lagenosoma propinquum Brauer, male; F. Lagenosoma propinquum Brauer, female; G. Lagenosoma varigatum sp. n., male; H. Lagenosoma varigatum sp. n., female; I. Lagenosoma propius sp. n., male.

FIGURE 37. Distribution of Lagenosoma species.