Stemonitis splendens var. flaccida Lister, Monogr.

Lado, Carlos & Basanta, Diana Wrigley De, 2018, Typification of the myxomycete taxa described by the Listers and preserved at the Natural History Museum, London (BM), Phytotaxa 341 (1), pp. 448-450 : 448-450

Stemonitis splendens var. flaccida Lister, Monogr. mycetozoa, ed. 1, 112. 1894

≡ Comatricha flaccida (Lister) Morgan, J. Cincinnati Soc. Nat.Hist. 16(4): 135. 1894

≡ Symphytocarpus flaccidus (Lister) Ing & Nann. -Bremek, Proc. Kon. Ned. Akad. Wetensch., C. 70(2): 217. 1967

— Lectotypus (designated here): U.S.A. Ohio, New Haven (Preston), 39º16’N 84º44’W, 1893, A.P. Morgan ( BM 1376 !, BM001089815 + one slide L:B.M.83!, engraved 301, BM001089168 ) . Syntypus: UNITED KINGDOM. England, Dorset, Lyme Regis , 50.725ºN 2.940ºW, Jun 1893, A. Lister (one slide L:B.M.83!, engraved 299, BM001089169 ) . Syntypus: GERMANY. Baden-Württemberg, Black Forest , 48º18’N 8º09’E, 18 Sep 1893, J.J. Lister (one slide L:B.M.83!, engraved 300, BM001089170 ) .

[BM] 1376 / Stemonitis splendens Rost. / γ. flaccida/ Preston, Ohio, U.S.A. / leg. A.P. Morgan 1893/ BM001089815 + one slide L:B.M.83 engraved 301/ BM001089168. L:B.M.83 one slide engraved 299/ Stemonitis / splendens Rost. / γ. flaccida/ Lyme Regis, Dorset/ leg. Lister June 1893 / BM001089169. L:B.M.83 one slide engraved 300/ Stemonitis / splendens Rost. / γ. flaccida/ Black Forest, Germany / leg. J. J. Lister Sep.18.93/ BM001089170.

NOTES: Lister (1894: 113–114) made reference to the following collections “Lyme Regis, Dorset (L:B.M.83); Black Forest (L:B.M.83); Ohio (L:B.M.83). In BM the slides of these three samples were located. The one from Ohio has a box (BM 1376) from the same locality, date and collector and we believe this is original material from which probably slide L:B.M.83 was taken. For this reason this sample was designated as the lectotype. There is a second box, BM 1377, that according to Ing & Nannenga-Bremekamp (1967), is the type for Symphytocarpus flaccidus (Lister) Ing & Nann. -Brem. and the generitypus specimen for the genus Symphytocarpus Ing & Nann. -Bremek The information on the label is: “[BM] 1377/ Stemonitis splendens Rost. / γ. flaccida/ on fir stumps/ Type (written in pencil)/ Shapwick, Lyme Regis/ leg. Lister July 6.1891” (with a revision label: “ Symphytocarpus flaccidus (Morg) / B. Ing & Nann.-Brem. nov. comb./ Type.– Koninkl. Nederl. Akad./ Wetensch. Ser. C, 70, 218. 1967”). However the collection date (July 6, 1891) does not coincide with that of the slide L:B.M.83 published by Lister (1894), that is dated June 1893, and so we do not believe it to be the original material.

We found comments in notebook IV: 71–73 on the material from Lyme Regis and in notebook VIII: 72 on Morgan’s American material. The var. flaccida , has the characters of “sporocarps weak, adhering to each other, the capillitium lax, scarcely forming a surface net, and the membranous flakes of sporocarps-wall always present”. These characters currently place it in the genus Symphytocarpus and it is the basionym of the species Symphytocarpus flaccidus (Lister) Ing & Nann. -Bremek

In K (http://apps.kew.org/herbtrack/results) one collection [K173669, England, 1850, A. Bloxam (ex herb. W.A. Leighton), A. Lister 715] is preserved but this material is from a previous date and is not cited in the publication.