Tambana Moore, 1882

Behounek, G., Han, H. L. & Kononenko, V. S., 2015, A revision of the genus Ta mb ana Moore, 1882 with description of eight new species and one subspecies (Lepidoptera, Noctuidae: Pantheinae). Revision of Pantheinae, contribution XIII, Zootaxa 4048 (3), pp. 301-351 : 303-304

Genus Tambana Moore, 1882

Tambana Moore, 1882 , in Hewitson & Moore, 1882, Description of new Indian Lepidopterous Insects from the Collection of the late Mr W. S. Atkinson (2): 155. Type species: Tambana variegata Moore, 1882 , by original designation. Synonymy: Trisulipsa Berio, 1973 , Annali del Museo Civico di Storia Naturale Giacomo Doria, 79: 160. Type species: Trisulipsa quadrata Berio, 1973 , by original designation. A junior synonym of Tambana bella ( Mell, 1935) . Trisuloides auct., nec Butler, 1881.

References. Warren 1912b: 43; Hampson 1913: 344; Prout 1922:227; Nye 1975: 470; Poole 1989:948, 978 (as synonym of Trisuloides , Trisulipsa Berio, 1973 , Trisuloides , part); Speidel & Kononenko 1998: 549 ( Tambana , syn. Trisulipsa , Trisuloides auct.).

Diagnosis. Tambana is a typical Pantheinae genus which could be characterized by the most often large size, hairy eyes, in most cases brown, grey-brown or grey colouration of forewing with characteristic wing pattern, yellow hindwing with dark terminal band (which sometimes covers the entire wing). Certain species have, however, white and black mosaic pattern or white-spotted dark-grey forewings, being similar to the members of Trichosea Grote, 1875 . The armature of the male genitalia in most cases has a rather uniform structure showing the affinity of Tambana with other Pantheinae genera, i. e. Trisuloides , Disepholcia , Trichosea , Xanthomantis , Antitrisuloides , and especially with Anacronicta Warren, 1913 , while the structure of the aedeagus and vesica, as well as the female genitalia, are rather diverse in the genus. The closest relative of Tambana is the genus Anacronicta (presently under revision), which in the most cases have similar with Tambana structural external characters, type of forewing pattern and shape or armature genitalia. Most distinctive character between Tambana and Anacronicta is structure of the vesica of the later, bearing by 2-3 large finger-like diverticula armed with dense medium cornuti. The structure of vesica in Tambana is variable, but without clearly expressed finger-like extensions.

Imago ( Figs 1–88 ). Medium-sized to large moths, wingspan 32–65 mm. Male antennae bipectinate ( T. albiplaga and T. plumbea ), serrate or filiform in other species; those of female filiform with fasciculate cilia; proboscis developed, short; labial palps short, compressed, 3rd segment short or moderate; eyes covered with short hairs; body stout; abdomen with crest formed by erect scales on 1–5 segments; forewing broad, usually dark brown, brown or grey; in most cases wing pattern with characteristic shape of subterminal line and tornal mark; hindwing colouration in most cases yellowish, yellow or orange with dark terminal band, in some species uniformly yellow, grey-brown or white.

Male genitalia ( Figs 89–117 ). Uncus rather short, broad, belt-like, beak or diamond-like, with pointer apex; tegumen broad, in some species with penicular lobes, vinculum large, V-shaped; juxta plate-like; transtilla with broad lobes; valva lobe-like, apically rounded; costa strong, sacculus narrow, harpe large, moderate curved, placed in mid or in distal third of part of valva. Aedeagus variable in shape, in most cases broad, massive; vesica tubular or bulbous-tubular, in some species very short; arming of vesica variable, in some species vesica unarmed.

Female genitalia ( Figs 118–130 ). Ovipositor with papillae anales short, broad, anterior and posterior apophyses relatively short, equal in length; antrum in many cases deep cup-like; ductus bursae moderately long, in some species sclerotised; corpus bursae large, saccular, rounded or elongate, often with sclerotised patch in caudal part.

The genus includes now 26 distinguished taxa (eight new species and a new subspecies are described here) distributed almost exclusively in the Oriental region, mainly in South and Southwest China and Indochina. Most species occur in montane, predominantly primary, forests.

Figures

FIGURES 1 – 8. Tambana spp., adults. 1. T. albiplaga, male, lectotype, N India (NHM); 2. T. albiplaga, female, Thailand (AB / ZSM); 3. T. arctoides, male, holotype, North Vietnam, slide VK 1695 (ZFMK); 4. T. arctoides, female, Myanmar (AB / ZSM); 5. T. bella, male, lectotype, China, Guanxi (ZFMK); 6. T. bella, male, paralectotype of T. chekiana, China, Zhejiang (ZFMK); 7. T. bella, male, holotype of T. quadrata, Myanmar (NHRM); 8. T. bella, male, China, Guangdong, slide HHL- 1502 - 1 (NEFU).

FIGURES 9 – 16. Tambana spp., adults. 9. T. burmana, male, holotype, Myanmar (NHRM); 10. T. burmana, male, China, Fujian (PGM); 11. T. burmana, female, China, Sichuan, slide GB 12122 (PGM); 12. T. burmana, female, China, Sichuan, slide GB 12130 (PGM); 13. T. naumanni, male, holotype, China, Guangdong (ZFMK); 14. T. naumanni, male, paratype, China, Guangdong (ZFMK); 15. T. plumbea, female, holotype, Japan, Tokyo (NHM); 16. T. plumbea, male, Russia, Sakhalin (GR).

FIGURES 17 – 24. Tambana spp., adults. 17. T. subflava, male, holotype, Taiwan (NHM); 18. T. subflava, female, holotype of T. albitessellata, N India (NHM); 19. T. subflava, male, South China, slide VK 1875 ZFMK (ZFMK); 20. T. subflava, female, China, Jiangxi / Fujian, slide GB 12053 (AZ / ZFMK); 21. T. subflava, male, Taiwan (GR); 22. T. tibetica sp. n., male, holotype, China, Tibet, slide GB / ZSM N 4290 (ZSM); 23. T. succincta, male, holotype, Myanmar (NHRM); 24. T. succincta, male, Myanmar, slide GB 12162 (AB / ZSM).

FIGURES 25 – 32. Tambana spp., adults. 25. T. variegata, male, holotype N India (NHM); 26. T. variegata, male, Myanmar (NHRM); 27. T. variegata, male, North Vietnam (GR); 28. T. entoxantha, male, syntype North India, Sikkim (NHM); 29. T. entoxantha, male, holotype of klapperichii, China, Guangdong (ZFMK); 30. T. entoxantha, female, holotype China, Jianxi / Fujian (AB / ZSM); 31. T. c-album, female, syntype China (NHM); 32. T. c-album, male, China, Hubei (GR).

FIGURES 33 – 40. Tambana spp., adults. 33. T. c-album, male, China, Shaanxi, slide GB 8242 (GB / ZSM); 34. T. c-album, male, China, Shaanxi, slide GB 12166 (JS / ZSM); 35. T. similina, male, holotype, China, Yunnan slide, DS-ZFMK (ZFMK); 36. T. similina, male, paratype, China, Yunnan, slide VK 2018 ZFMK (ZFMK); 37. T. nekrasovi, male, holotype, China, Shaanxi, slide VK-ZISP (ZISP); 38. T. nekrasovi, male, China, Sichuan, slide GB 1218 (PGM); 39. T. xilinga sp. n., male, holotype, China, Sichuan, slide GB 7401 (GB / ZSM); 40. T. sp., male, Myanmar, slide GB 7401 (GB / ZSM).

FIGURES 41 – 46. Tambana spp., adults. 41. T. mekonga sp. n., male, holotype, China, Yunnan, slide GB 12074 (GB / ZSM); 42. T. mekonga sp. n., male, paratype, China, Yunnan, slide GB 8676 (GB / ZSM); 43. T. helmuti, male, holotype, Myanmar, slide GB 12020 (GB / ZSM); 44. T. fansipana sp. n., female, holotype, Vietnam, slide GB 7402 (GB / ZSM); 45. T. funebris, male, holotype, Myanmar (NHRM); 46. T. funebris, male, paratype of T. gerri, Vietnam, slide GB 8050 (GB / ZSM); 47. T. funebris, female, China, Sichuan, slide GB 12106 (PGM); 48. T. ronnyi, male, paratype, Vietnam, slide GB 7707 (GB / ZSM).

FIGURES 49 – 56. Tambana spp., adults. 49. T. laura sp. n., male, holotype, China, Yunnan, slide GB 5826 (AB / ZSM); 50. T. laura sp. n., male, paratype, Myanmar, slide GB 12005 (HS); 51. T. glauca, North India, holotype, slide MH 12029 / Noctuidae Brit. Mus. Slide No 15806 (NHM); 52. T. glauca, male, holotype of T. behouneki North Vietnam, slide GB 5375 (GB / ZSM); 53. T. glauca, male, Thailand, slide GB 7404 (GB / ZSM); 54. T. glauca, male, Thailand, slide GB 7407 (GB / ZSM); 55. T. glauca, male, Thailand, slide GB 7423 (AB / ZSM); 56. T. glauca, female, Myanmar (AB / ZSM).

FIGURES 57 – 64. Tambana spp., adults. 57. T. indeterminata sp. n., male, North India, holotype, slide GB 7403 (GB / ZSM); 58. T. indeterminata sp. n., male, paratype, China, Yunnan, slide GB 12145 (WS / ZSM); 59. T. indeterminata sp. n., male, paratype, China, Xizang, slide HHL- 1773 (NEFU); 60. T. indeterminata sp. n., male, paratype, North India, slide GB 12164 (JS); 61. T. annamica, male sp. n., holotype, North Vietnam, slide GB 7405 (GB / ZSM); 62. T. annamica, male sp. n., paratype, North Vietnam, slide GB 7406 (GB / ZSM); 63. T. annamica sp. n., male, paratype, North Vietnam, slide RL 6669 (HNHM); 64. T. annamica sp. n., male, paratype, North Vietnam (AB / ZSM).

FIGURES 65 – 88. Tambana spp., adults. 65. T. annamica stumpfi ssp. n., male, holotype, China, Guangxi, slide GB 12113 (PGM); 66. T. annamica stumpfi ssp. n., male, paratype, China, Shaanxi (JS); 67. T. annamica stumpfi ssp. n., male, paratype, China, Guangxi, slide GB 12112 (PGM); 68. T. annamica stumpfi ssp. n., male, paratype China, Guangdong, slide HHL- 1500 - 1 (NEFU); 69 – 73. Series of T. glauca sp. n. (North India, North Vietnam, North Thailand, Myanmar, Southwest China); 74 – 78. Series of T. indeterminata sp. n. (North India, North Thailand, Southwest China); 79 – 83. Series of T. annamica sp. n. (North Vietnam); 84 – 88. Series of T. annamica stumpfi ssp. n. (China: Shaanxi, Guangxi, Guangdong).

FIGURES 89 – 91. Tambana spp., male genitalia. 89. T. albiplaga, North Vietnam, slide WS 377 (WS); 90. T. arctoides, holotype, North Vietnam slide VK 1656 (ZFMK); 91. T. bella, China, Guangdong, slide HHL- 1502 - 1 (NEFU).

FIGURES 92 – 94. Tambana spp., male genitalia. 92. T. burmana, paratype, Myanmar, slide VK / NHRM- 2 (NHRM); 93. T. naumanni, paratype, China, Guangdong, slide VK 1784 (ZFMK); 94. T. plumbea, Russia, Sakhalin, slide VK 277 (IBSS).

FIGURES 95 – 97. Tambana spp., male genitalia. 95. T. subflava, China, Yunnan, slide HHL- 1770 - 1 (NEFU); 96. T. tibetica, China, Tibet, slide GB ZSM N 4290 (ZSM); 97. T. succincta, paratype, Myanmar slide VK / NHRM- 2 (NHRM).

FIGURES 98 – 100. Tambana spp., male genitalia. 98. T. variegata, North Vietnam, slide WS 378 (WS); 99. T. entoxantha, Indonesia, Sumatra, slide AZ 12049 - 1 (AZ); 100. T. c-album, China, Shaanxi, slide VK 2018 (ZFMK).

FIGURES 101 – 103. Tambana spp., male genitalia. 101. T. similina, holotype, China, Yunnan, slide DS- 2013 (ZFMK); 102. T. nekrasovi, holotype, China, Shaanxi, slide VK-ZISP, coll. Nekrasov / ZISP; 103. T. xilinga sp. n., holotype, China, Sichuan, slide GB 8066 (GB / ZSM).

FIGURES 104 – 106. Tambana spp., male genitalia. 104. T. sp., Myanmar, slide GB 7401 (AB / ZSM); 105. T. mekonga sp. n., holotype, China, slide GB 12074 (WS / ZSM); 106. T. helmuti sp. n., holotype, Myanmar, slide GB 12020 (HS / NHM, Vienna).

FIGURES 107 – 109. Tambana spp., male genitalia. 107. T. funebris, paratype, Myanmar, slide VK / NHRM- 2 (NHRM); 108. T. ronnyi, paratype, Vietnam, slide GB 7707 (GB / ZSM); 109. T. laura sp. n., holotype, China, Yunnan, slide GB 5826 (AB / ZSM).

FIGURES 110 – 113. Tambana spp., male genitalia. 110. T. glauca, North Vietnam, holotype of behouneki, slide 5375 (WS / ZFMK); 111. T. glauca, Thailand, slide GB 7423 (GB / ZFMK); 112. T. indeterminata sp. n., holotype, North India, slide GB 7403 (GB / ZSM); 113. T. indeterminata sp. n., paratype, China, Guangxi slide HHL- 2606 - 17403 (NEFU).

FIGURES 114 – 117. Tambana spp, male genitalia. 114. T. annamica sp. n., holotype, North Vietnam, slide GB 7405 (GB / ZSM); 115. T. annamica sp. n., paratype, North Vietnam, slide 6669 (HNHM); 116. T. annamica stumpfi ssp. n., paratype, China, Guangxi, slide GB 12113 (PGM); 117. T. a. stumpfi ssp. n., China, Guangdong, paratype, slide HHL- 1500 - 1 (NEFU).

FIGURES 118 – 123. Tambana spp., female genitalia. 118. T. albiplaga, North Vietnam, slide GB 8254 (GB / ZSM); 119. T. arctoides, Myanmar, slide GB 12161 (AB); 120. T. bella, China, Yunnan, slide GB 12117 (PGM); 121. T. fansipana sp. n., holotype, North Vietnam, slide GB 7402 (GB / ZSM); 122. T. burmana, China, Sichuan, slide GB 12122 (PGM); 123. T. naumanni, paratype, China, Guangdong, WS 412 (ZFMK).

FIGURES 124 – 130. Tambana spp., female genitalia. 124. T. plumbea, Russia, Sakhalin, slide VK 276 (IBSS); 125. T. subflava, China, Yunnan, slide GB 12053 (AZ); 126. T. c-album, China, Shaanxi, slide GB 12167 (JS); 127. T. entoxantha, Indonesia, Sumatra, slide GB 7704 (GB / ZSM); 128. T. funebris, China, Sichuan, slide 12106 (PGM); 129. T. glauca, paratype of behouneki, North Vietnam, slide WS 413 (WS); 130. T. glauca, Thailand, slide GB 7404 (GB / ZSM).