Liquidambar europaea A. BRAUN

Kovar-Eder, Johanna, Kvaček, Zlatko, Teodoridis, Vasilis, Mazouch, Petr & Collinson, Margaret E., 2022, Floristic, Vegetation And Climate Assessment Of The Early / Middle Miocene Parschlug Flora Indicates A Distinctly Seasonal Climate, Fossil Imprint 78 (1), pp. 80-144 : 96-97

Liquidambar europaea A. BRAUN

2004 Liquidambar europaea A.BRAUN ; Kovar-Eder et al., p. 58, pl. 2, figs 1–5.

A d d i t i o n a l m a t e r i a l. GBA 1848/0001/0050, GBA 2002/0001/0094a, GBA 2005/0004/0090, 0109, 0114A, IBUG 1185, NHMW 1878/6/2336.

D e s c r i p t i o n. Leaves long petiolate; trilobate to double quinquelobate; base shape lobate; l (central lobe) × w (widest distance between lobe tips) up to at least 90 × 138 mm; medial and inner lateral lobes ovate to often obovate or slender elongated, rarely with additional two to four smaller lobes ( form often designated as Liquidambar parschlugiana UNGER ); outermost lateral lobes smaller, narrow triangular; lobe apices attenuate to bluntly acute, sinuses between lobes acute; margin regularly, finely glandular serrate; venation palmate; midvein strong, straight; 2–4 lateral primaries arising from the base, straight to slightly bent towards base; secondaries semicraspedodromous (and craspedodromous in double lobed leaves), widely regularly spaced, curved towards apex (convex); tertiaries irregular, forming elongated meshes oblique to the secondaries.

Mahonia (?) sphenophylla (UNGER) DOWELD Pl. 7, Figs 12–16

2004 Mahonia (?) aspera (UNGER) KOVAR- EDER et KVAČEK; Kovar-Eder et al., p. 57, pl. 13, figs 1–8.

A d d i t i o n a l m a t e r i a l.GBA1848/0001/0086, GBA 2005/0004/cf. 0001, 0034, cf. 0051, NHMW 1878/6/2031, 2381, 2406a, 9497, Ett. 642c.

D e s c r i p t i o n. Leaflets sessile or shortly petiolulate; lamina coriaceous, shape roundish to obovate, l × w about 12–48 (85) × 8–20 (30) mm, ratio l/w about 1–2.4, base subcordate to convex, sometimes slightly asymmetric, apex rounded; margin irregularly widely simple dentate to entire, teeth spine-like; venation acrodromous; midvein thick, straight; lateral primaries basal or shortly suprabasal, gently bent, reaching to about half of the lamina length; secondaries semicraspedodromous in 2–3 pairs, looping with lateral primaries, in dentate forms sending side veinlets into teeth; intersecondaries occasionally present; in the lower part tertiaries connecting midvein with lateral primaries percurrent, otherwise tertiaries reticulate, forming polygonal irregular meshes, rarely visible due to coriaceous texture of leaves.

R e m a r k s. Doweld (2018a) recognised that the combination Quercus aspera had been invalidly published by Unger (1847). Therefore, the oldest available name for this taxon is Ilex sphenophylla also published by Unger (1847) (see also synonymy for Mahonia (?) aspera in Kovar-Eder et al. 2004).