Celastrus

Kovar-Eder, Johanna, Kvaček, Zlatko, Teodoridis, Vasilis, Mazouch, Petr & Collinson, Margaret E., 2022, Floristic, Vegetation And Climate Assessment Of The Early / Middle Miocene Parschlug Flora Indicates A Distinctly Seasonal Climate, Fossil Imprint 78 (1), pp. 80-144 : 87

“ Celastrus ” europaea UNGER

Pl. 1, Figs 10–12

2004 “ Celastrus ” europaea UNGER ; Kovar-Eder et al., p. 84, pl. 12, figs 1–2.

2004 “ Euonymus ” latoniae UNGER; Kovar-Eder et al., p. 85, pl. 12, figs 3, 5 (non fig. 4).

A d d i t i o n a l m a t e r i a l. IBUG 1988, NHMW 1878/6/cf. 2740 + 2741 (part + counterpart), 2742.

D e s c r i p t i o n. Leaves (?) short-petiolate; lamina elliptic, l × w about 31–46 × 17–21 mm, ratio l/w = 1.8–2.7, base cuneate/decurrent, apex bluntly acute, tip rounded; margin basally entire, in the upper part almost entire/ slightly undulate to toothed with very few and more or less distinct blunt teeth; midvein straight; secondaries (?) eucamptodromous to semicraspedodromous, widely spaced, arising at acute angles, rather straight; intersecondaries occasionally present; tertiaries reticulate.

R e m a r k s. After studying more material, we merged both morphospecies because we are unable to differentiate them. Hably (2020) included the leaves published as “ Euonymus ” latoniae by Kovar-Eder et al. (2004) into synonymy with cf. Gleditsia sp. The leaves from Parschlug differ, however, from cf. Gleditsia sp. by their symmetrical shape, the presence of a petiole and blunt teeth. Only specimen NHMW 1878/6/2063 ( Kovar-Eder et al. 2004: pl. 12, fig. 4), transferred here to Ailanthus pythii , resembles the specimen figured by Hably (2020) on pl. 17, fig. 4 in shape and the widely, irregularly spaced, blunt teeth.

Differing from Dicotylophyllum sp. B by non-entire margin, more widely and regularly spaced, steeper secondaries and short petiole.

Differing from Dicotylophyllum sp. X by non-entire margin.

Differing from Dicotylophyllum sp. HH by toothed margin and absence of a long petiole.

Differing from Toxicodendron melaenum by symmetrical shape and smaller, often indistinct teeth.