Adelostoma (Zarudnionymus) abyssinicum obockensis, Purchart, Luboš, 2012

Purchart, Luboš, 2012, Revision of the genus Adelostoma (Coleoptera: Tenebrionidae). Part 1: subgenus Zarudnionymus Semenov & Bogatchev, 1947, Zootaxa 3164, pp. 17-31 : 24-25

Adelostoma (Zarudnionymus) abyssinicum obockensis subsp. nov.

( Fig. 3 , 10 , 17 , 24 )

Type locality. Djibouti, Obock.

Type material. Holotype (1 3 MNHN): Obock, Dr. Martin {printed} // MUSEUM PARIS, 1922, Coll. L. BEDEL .

Paratypes (7 MNHN, 1 LPCB): same data as holotype ; (1 3 MNHN): Obock {handwritten} // MUSEUM PARIS, COLL. P. ARDOIN, 1978 {blue label, printed} ; (2 HNHM): Obock // COLL. P. DE PEYERIMHOFF ; (2 MNHN): COLL. P. DE PEYERIMHOFF {blue label, printed} // Abyssinium, Obock, leg. Théry {handwritten} ; (5 MNHN, 1 LPCB): Obock, Coll. Sietti {printed}// MUSÉUM PARIS, 1949, Coll. H. SIETTI {blue label, printed} ; (4 MNHN, 1 LPCB): Obock {handwritten} // MUSÉUM PARIS, 1898, COLL. A. CHOBAUT {blue label, printed} ; (2 MNHN): Obock, ... {unreadable, handwritten} // MUSEUM PARIS, Collection Léon Fairmaire, 1906 ; (1 MNHN): MUSEUM PARIS, DJIBOUTI, JOUSSEAUME [leg.], 7-97 [1897] ; (1 MNHN): MUSEUM PARIS, DJIBOUTI, H. COUTIÈRE [leg.], 1897 ; (1 MNHN): MUSEUM PARIS, OBOK [=Obock], JOUSSEAUME [leg.], 7-97 [1897] ; (3 MNHN): Obock, Dr. Martin // MUSÉUM PARIS, 1937, COLL. C. DEMAISON ; (1 MNHN): Obock, Jousseuame [leg.] {handwritten} // abyssinicum Haag {handwritten} // MUSEUM PARIS, 1922, Coll. K. BEDEL ; (1 MNHN): Adelostoma abyssinicum {handwritten} // MUSEUM PARIS, 1922, Coll. K. BEDEL ; (1 MNHN): no locality data // MUSEUM PARIS, 1922, Coll. K. BEDEL ; (2 MNHN): Obock, Dr. J. Besnard ; (1 MNHN): Obock, Dr. Martin // Museum Paris, Coll. M. Pic ; (5 BMNH): 2244 {rounded blue, handwritten} // without locality labels.

Differential diagnosis. A. a. obockensis subsp. nov. can be distinguished from A. borowieci and A. grande by short third antennomere and with more or less convex pronotum ( Fig. 10 , 17 ). In A. borowieci and A. grande the third antennomere is prolonged ( Fig. 19, 21 ) and shape of pronotum different—cordiform in A. grande ( Fig. 14 ), narrow, not transverse and nearly square in A. borowieci ( Fig. 12 ). From A. batesi it differs by complete median carina of head and shape of pronotum. In A. batesi median carina of head is short, not complete and pronotum is strongly transverse and cordiform ( Fig. 11 ). From A. gilloni it can be separated by the shape of pronotum, by short third antennomere and by elytral keels which are well developed, not obliterated. In A. gilloni the pronotum is distinctly cordiform ( Fig. 13 ), the third antennomere prolonged ( Fig. 20 ) and at least third of the outer keel of elytra is obliterated. The new subspecies most resembles A. a. abyssinicum and A. a. hirsutum . They differ in shape of pronotum, which is distinctly cordifom with slightly and irregularly serrate or dentate ( Fig. 8, 9 ) lateral margins, in contrast to A. a. obockensis in which the pronotum is not cordiform but more or less convex with distinctly and regularly dentate lateral margins ( Fig. 10 ). A. a. obockensis differs also in the following characters: reflected portion of elytra (sensu Koch 1952: 5, 7) is broader and in the level of 1st and 2nd abdominal ventrites with at least four (often five) puncture intervals and with punctures arranged more or less regularly in rows ( Fig. 24 ). In A. a. hirsutum 3–4 puncture intervals are present. Punctures are arranged irregularly ( Fig. 23 ). In A. a. abyssinicum three punctures intervals can be observed. Punctures are arranged in regular longitudinal rows ( Fig. 22 ).

Description. Holotype, body length 6.2 mm, width 2.5 mm. Body narrow, parallel. Dorsal and ventral side of body brown and dull.

Head 0.87 times shorter than broad, 0.75 times narrower than pronotum. Widest at genae. Shagreened, covered sparsely with whitish hairs, denser on epistome. Length of hairs approximately as long as length of second antennomere. Surface of head sculptured with net-like surface composed of concave granules which approximately 2–3 times larger than one eye facet. Genae strongly developed. Ocular carinae high (lateral view). Midlongitudinal carina complete, starting at level of posterior part of eyes and reaching anterior margin of clypeus, lower than ocular carinae (lateral view). Depressed area between antennal sulcus and midlongitudinal carina large and deep. Clypeus asymmetrically cut out in the middle and with three small teeth in the right part of the latter (this character seems to be variable as number of teeth in paratypes varying between 1–3). Antennae 1.33 times longer than width of head, covered with whitish setae. Antennomeres globose, with obliquely truncate apical antennomere, which is longer than broad, approximately two times longer than penultimate antennomere.

Pronotum transverse, 1.43 times wider than long, more or less convex, widest in middle. Shagreened, sparsely covered with recumbent and relatively long yellowish hairs. Sculptured with net-like surface composed of irregular concave granules which approximately 4–6 times larger than one eye facet. Lateral portions of pronotum foliaceous, flattened. Two parallel carinae well developed and complete. Anterior and posterior angles of pronotum extended to spikes. Lateral margins of pronotum distinctly and more or less regularly dentate.

Elytra shagreened, parallel, with three longitudinal, strongly developed and relatively high keels on each elytron. Space between keels irregularly deeply punctuate with 3–4 puncture intervals between each keel. Surface between these punctures sometimes with very small tubercles smaller than one eye facet. First (inner, not sutural) and third (outer) keels connected anteriorly with basal margin of elytra. Third and second (median) keels not connected with apex of elytra. First carina almost reaching apex of elytra and connected with epipleural carina. The latter developed only apically. Reflected portion of elytra (sensu Koch 1952: 5, 7) at level of 1st and 2nd abdominal ventrites with four puncture intervals, with punctures arranged more or less regularly in rows. Entire surface of elytra sparsely covered with same hairs as pronotum and with protruding and long yellowish hairs especially at apical part of elytra. Apex of elytra moderately steep (lateral view).

Legs shagreened and covered with brownish setae, without any special features.

Entire ventral side of head and prothorax sculptured with net-like surface composed of concave granules and shagreened, covered with hairs similar to those on dorsal side of head. The rest of ventral side of body shagreened, deeply punctured, sparsely covered with short whitish hairs. Epipleural carina developed only apically.

Aedeagus simple.

Variability in size: 5.6–6.5 × 2.0–2.6 mm.

Distribution. Djibouti.

Figures

FIGURES 1 – 7. Habitus, dorsal view. 1. Adelostoma abyssinicum abyssinicum (non-type); 2. A. a. hirsutum (non-type); 3. A. a. obockensis subsp. nov. (holotype); 4. A. batesi (non-type); 5. A. borowieci (non-type); 6. A. gilloni (non-type); 7. A. grande (non-type).

FIGURES 8 – 14. Pronotum, dorsal view. 8. Adelostoma abyssinicum abyssinicum (non-type); 9. A. a. hirsutum (non-type); 10. A. a. obockensis subsp. nov. (holotype); 11. A. batesi (non-type); 12. A. borowieci (non-type); 13. A. gilloni (non-type); 14. A. grande (non-type).

FIGURES 15 – 21. Antenna, dorsal view. Arrows indicate elongate third antennomere. 15. Adelostoma abyssinicum abyssinicum (non-type); 16. A. a. hirsutum (non-type); 17. A. a. obockensis subsp. nov. (holotype); 18. A. batesi (non-type); 19. A. borowieci (non-type); 20. A. gilloni (non-type); 21. A. grande (non-type).

FIGURES 22 – 24. Reflected portion of elytra (RFPE) with punctured intervals. Rectangles indicate the area first (1 st) and second (2 st) abdominal ventrites. Arrows indicate the range of RFPE. Latero-ventral view. 22. Adelostoma abyssinicum abyssinicum (non-type); 23. A. a. hirsutum (non-type); 24. A. a. obockensis subsp. nov. (paratype).