Ocypus Leach, 1819

Zhou, Hong-Zhang, 2017, Taxonomy of the subgenus Ocypus Leach (Coleoptera: Staphylinidae: Staphylinini) and three new species from China, Zootaxa 4312 (2), pp. 293-310 : 294

Subgenus Ocypus Leach, 1819

Leach, 1819: 172 (species included: cyaneus ); Bernhauer and Schubert, 1914: 374 (subgenus of Staphylinus ; authorship as Stephens, 1832; world catalog); Cameron, 1932: 180 (subgenus of Staphylinus ); Scheerpeltz, 1933: 1381 (subgenus of Staphylinus , catalog supplement); Shibata, 1973: 62 (checklist of species of Taiwan); Coiffait, 1974: 420 (key to species of the western Palearctic Region); Shibata, 1984: 87 (checklist of species of Japan); Smetana and Davies, 2000: 21, 31, 42 (characters; checklist of species of north temperate region); Herman, 2001: 3355 (world catalog); Smetana, 2004: 674 (Catalog of Palaearctic Region); Smetana, 2007: 443 (key to species of China); Schülke & Smetana, 2015: 1089 (Catalog of Palaearctic Region, including the synonyms Nudabemus and Xanthocypus).

Type species: Staphylinus cyaneus Paykull, 1789 , fixed by original designation and monotypy. Syn.: Goerius Westwood, 1827: 58 . Type species: Staphylinus olens O. Müller, 1764 by monotypy. Isopterum Gistel, 1856: 388 . Type species: Staphylinus cyaneus Paykull, 1789 , fixed by subsequent designation by

Blackwelder, 1952: 2015.

Nudabemus Coiffait, 1982b: 74 . Type species Nudabemus caerulescens Coiffait, 1982 , fixed by original designation. Xanthocypus J. Müller, 1925d: 40. Type species Ocypus weisei Harold, 1877 .

The species of the subgenus Ocypus s. str. can be easily distinguished from those of other subgenera of the genus Ocypus by the following characteristics: last segment of maxillary palpi asetose; that of labial palpi setose, usually short, with truncate apex; body size relatively large (more than 15 mm), body color usually totally black; palpifer usually with three setae on apical portion; eyes small and tempora relatively long; genal seta at least four puncture diameters away from posterior margin of eye; sternite 9 of male genital segment with tapered basal portion rather short and wide.

In the current taxonomy ( Smetana and Davies, 2000; Herman, 2001; Smetana 2007; Schülke & Smetana, 2015), the subgenus Ocypus s. str. includes the synonyms Goerius Westwood, 1827 , Isopterum Gistel, 1856 , Nudabemus Coiffait, 1982 , and Xanthocypus J. Müller, 1925. The taxonomic position of the name Xanthocypus was once challenged if it should be maintained within the subgenus Ocypus s. str. ( Hayashi, 1993; Smetana 2007: “tentatively set aside as a possible junior synonym of Agelosus Sharp, 1889 ”), but this was not confirmed by later publications (Smetana, 2009; Schülke & Smetana, 2015). We therefore maintained it within the subgenus Ocypus s. str. and included here two Chinese species: Ocypus miwai ( Bernhauer, 1943) and Ocypus weisei Harold, 1877 .