Metriorhynchus superciliosus, (BLAINVILLE, 1853), 1869

Young, Mark T., Brusatte, Stephen L., Ruta, Marcello & Andrade, Marco Brandalise De, 2010, The evolution of Metriorhynchoidea (mesoeucrocodylia, thalattosuchia): an integrated approach using geometric morphometrics, analysis of disparity, and biomechanics, Zoological Journal of the Linnean Society (Zool. J. Linn. Soc.) 158 (4), pp. 801-859 : 848-850

METRIORHYNCHUS SUPERCILIOSUS (BLAINVILLE, 1853) EUDES- DESLONGCHAMPS, 1869

1824 ‘Gavial de Honfleur’; Cuvier [partim]

1824 ‘Deuxième gavial de Honfleur’; Cuvier [partim]

1825 Steneosaurus rostro -minor sp. nov.; Geoffroy Saint-Hilaire [partim]

1832 Metriorhynchus geoffroyii sp. nov.; von Meyer [partim]

1853 Crocodilus superciliosus sp. nov.; Blainville

1868b Teleosaurus superciliosus comb. nov.; Eudes- Deslongchamps

1868b Teleosaurus blainvillei sp. nov.; Eudes- Deslongchamps

1869 Metriorhynchus superciliosus comb. nov.; Eudes- Deslongchamps

1869 Metriorhynchus blainvillei comb. nov.; Eudes- Deslongchamps

1867–1969 Metriorhynchus moreli sp. nov.; Eudes- Deslongchamps

1869 Steneosaurus dasyceps sp. nov.; Seeley

1890b Metriorhynchus superciliosum unjust. emend.; Lydekker

1904 Metriorhynchus jaekeli sp. nov.; Schmidt

Neotype: MNHN 8903, incomplete skull (chosen by Blainville, see Vignaud, 1995).

Type locality: The original holotype is believed to be from England, see Vignaud (1995) for a full discussion (Oxford Clay Formation) .

Geological range: Lower Callovian–lower Oxfordian (koenigi–mariae ammonite zones).

Geographical range: Cambridgeshire, Huntingdonshire, and Wiltshire, England; and Ardèche , Normandy, and Poitou, France .

Geographical range note: Metriorhynchus specimens are known from the Middle Callovian of Germany (C. Diedrich, pers. comm., 2008).

Referred specimens: AMNH FR 997 , incomplete skull with mandible, and 23 assorted vertebrae (including cervicals and dorsals) ; CAMSM J64398 , skull (holotype of Steneosaurus dasyceps ) ; CAMSM J64900 , midportion of skull ; CAMSM J64918 , rostrum ; GLAHM V942 , skull with mandible from a young individual ; GLAHM V963 , incomplete skull and mandible, eight dorsal vertebrae, one caudal vertebra, both coracoids, left scapula, right ilium, left ischium, and right ischium blade, distal half of the right pubis (from a baby specimen) ; GLAHM V964 , skull, mandible, atlas axis, four cervical vertebrae, and cervical and dorsal ribs ; GLAHM V965 , skull with right dentary ; GLAHM V971 , skull in various fragments, atlas axis, five cervical vertebrae, and ten dorsal vertebrae ; GLAHM V982 , broken skull, mandible, cervical vertebra centrum, and atlas left rib ; GLAHM V983 , broken skull, mandible, sclerotic ring ossicles, and dorsal rib ; GLAHM V984 , broken skull and mandible fragments from two individuals ; GLAHM V985 , broken skull and mandible fragments, and sclerotic ring ossicles ; GLAHM V987 , broken skull and mandible fragments, sclerotic ring ossicles, and rib fragments ; GLAHM V988 , skull, mandible, atlas axis, one dorsal vertebra, left ilium, fragment of right pubis, and right femur ; GLAHM V989 , skull with mandible ; GLAHM V996 , skull with bite marks consistent with that of a metriorhynchid ; GLAHM V1004 , skull and mandible fragments, and atlas axis ; GLAHM V1015 , incomplete skull, disarticulated mandible, atlas axis, right atlas rib, both coracoids, right ischium, femora, right tibia and fibula, and two coprolites ; GLAHM V1027 , incomplete skull and mandible, atlas axis, four cervical vertebrae (from a young individual) ; GLAHM V1140 , broken skull, mandible showing pathological damage, one hyoid, atlas axis, five cervical vertebrae, 14 dorsal vertebrae, scapulae, both coracoids, humeri, both radii, and femora ; GLAHM V1142 , skull, disarticulated mandible, one hyoid, atlas axis, right atlas rib, four cervical vertebrae, 19 dorsal vertebrae, two sacral vertebrae, four caudal vertebrae, pubes, ilia, ischia (fused), and femora ; GLAHM V1143 , skull broken in two, mandible, atlas ribs, five cervical vertebrae, four cervical ribs, 17 dorsal vertebrae, right scapula, right coracoid, right humerus (atlas axis and forelimb lost) ; LEICT G1301.1899 , skull, disarticulated mandibular rami, atlas axis, two cervical vertebrae, 15 dorsal vertebrae, two sacral vertebrae, six caudal vertebrae, numerous ribs, right pubis, ilia, ischia, femora, right radius, ulnae, left astragulus and calcaneum, and fused tarsals ; MGCL 9959 , skull, mandible, associated dorsal and caudal vertebrae, humerus, femur ; MGCL 9960 , skull, mandible, associated cervical, dorsal and caudal vertebrae, isolated ribs, humerus, pubis, ischium, and femur; NHM R.1530, incomplete skull with mandible, atlas-axis, five cervical vertebrae, 16 dorsal vertebrae, two sacral vertebrae, 35 caudal vertebrae, cervical and dorsal ribs, coracoid, scapula, humeri, ilia, ischia, pubes, femora, tibiae, fibulae, isolated pes bones, and numerous chevrons; NHM R.1529, skull and mandible (from a young individual); NHM R.1666, skull and mandible; NHM R.2030, incomplete skull with mandible; NHM R.2032, incomplete skull and mandible, atlas axis, four cervical vertebrae, 17 dorsal vertebrae, two sacral vertebrae, 37 caudal vertebrae, cervical and dorsal ribs, scapula, coracoids, humerus, ilium, ischium, and femur; NHM R.2033, fragmentary skull with mandible, atlas axis, five cervical vertebrae, 17 dorsal vertebrae, two sacral vertebrae, 29 caudal vertebrae, cervical and dorsal ribs, ilia (fused), ischia (fused), pubes, one tibia, one fibula, and two metatarsals; NHM R.2036, fragments of skull and mandible, and one cervical vertebra; NHM R.2041, incomplete skull; NHM R.2044, skull and mandible; NHM R.2049, incomplete skull and mandible, two dorsal vertebrae, one sacral vertebra, ilia, ischia, femora, tibiae, fibulae, and isolated bones of the pes; NHM R.2051, incomplete skull with mandible, atlas axis, four cervical vertebrae, 15 dorsal vertebrae, two sacral vertebrae, 32 caudal vertebrae, coracoid, scapula, ilia, ischia, pubes, femora, broken tibiae and fibulae, and isolated pes bones; NHM R.2053, incomplete skull; NHM R.2054, skull, mandible, atlas axis, four cervical vertebrae, 18 dorsal vertebrae, two sacral vertebrae, 28 caudal vertebrae, cervical and dorsal ribs, ilium, ischium, pubes, tibia, fibula, isolated pes bones, numerous chevrons; NHM R.2055, incomplete skull; NHM R.2056, fragments of skull and mandible (from a very young individual); NHM R.2058, skull with mandibular symphysis; NHM R.2065, skull fragments (from a young individual); NHM R.2069, fragments of skull and mandible, atlas axis and three cervical vertebrae (from a very young individual); NHM R.3016, incomplete skull, mandible, right coracoid, scapulae, humeri, radius, ulna; NHM R.3900, rostrum; NHM R.6859, skull with disarticulated mandible; NHM R.6860, skull with disarticulated mandible; RMS M150, incomplete skull with mandible; NMING F16892, skull with disarticulated mandible, and 16 associated vertebrae ; NMING F21731, skull with disarticulated mandible, 28 associate vertebrae, and ilia ; NMW 19.96 G21, skull with mandible ; PETMG R8 , skull lacking rostrum ; PETMG R10 , incomplete skull ; PETMG R20 , incomplete skull ; PETMG R42 , incomplete skull ; PETMG R180 , incomplete skull ; SMNS 10115 , skull with disarticulated mandible ; SMNS 10116 , skull with disarticulated mandible .

Note: The correct spelling is M. superciliosus , not M. superciliosum . Lydekker (1890b) amended the epithet based on the neuter suffix, rather than retaining the masculine suffix (i.e. - sum instead of - sus). In accordance with the code of the ICZN, Article 30.1.3, the gender of the generic and epithet name must match, and as Metriorhynchus is masculine (- rhynchus is the Latinized form of the Ancient Greek - rhynchos) - sus is the correct suffix.