Chelipoda rangopango, Published, 2007

Published, First, 2007, The Hemerodromiinae (Diptera: Empididae) of New Zealand II. Chelipoda Macquart, Zootaxa 1537 (1), pp. 1-88 : 52-53

34. Chelipoda rangopango sp. n.

[ Figs. 55 , 79]

Type material. Holotype ♂: New Zealand, South Is. , NC, Glynn Wye Station, Kakapo Brook Val., site code ‘ Kakapo’, 42° 39`S, 172° 23`E, FIT on ground in forest, 3–16/i/2001, RKD [ NZAC] . Paratypes: 1♂, 1♀ same data as holotype ; 1♂, 3♀ same data as holotype, on ground at forest edge, 6-20/xii/2000 [ NZAC, NMWC] ; 1♂ same data as holotype, site code ‘ Donut’, 42° 39`S, 172° 22`E, 1–15/i/2001 [ NMWC] and 1♀, FIT on ground in forest, 6–18/xii/2000 [ NZAC]: North Is .; 1♂, 3♀, WO, Mt. Karioi , Raglan, Malaise by Wainui Str., 5–11/x/ 1985, 12–20/x/1985, 7–14/ix/1986, HAO [ NMWC] ; 2♂, 5♀, TO, Urewera NP, W of pass on SH 38, 760m, 5/ ix/1977, ES [ NZAC] .

Etymology: from the Maori; rango, fly; pango, black.

Description. Male: length = 2.5–3.0mm.

Head: subspherical, black, dusted greyish, setae black; verticals about as strong as ocl, not contiguous with upo; lpo becoming longer and brownish below, diverging from eye margin; no fine pile on lower occiput. Frons slightly produced anteriorly above antennae; eyes narrowly separated on face; hypostomal bridge rather large forming flange at back of head below occipital foramen. Antennae black, basal segments and postpedicel basally yellowish below; scape 2X as long as wide, a distinct dorsal seta reaching to end of pedicel; postpedicel strongly dorsoventrally flattened, 1.5X as long as wide in dorsal view (3X as long as wide in lateral view), arista 1.3X as long. Mouthparts yellowish brown.

Thorax: (Fig. 79) black, all setae black and rather stout; dc1 very short but strong, 0.2X as long as dc2; dc2, dc3, unp, pprn and sa strong; dc4 and ph minute; lnp weaker than unp (sometimes still smaller bristle above and in front).

Legs: blackish; tibiae, tarsi basally, femora apically and C 1 apically yellow; C 2 and C 3 yellowish brown. C 1 0.9X as long as thorax, only small bristly hairs anteriorly; F 1 1.2X length of C 1, not strongly inflated, widest 0.3 from base narrowing gradually distally; an av series of 10–11 denticle-like spines; pv series of 6–7 distinct spines (often with a few much smaller spines intermingled) longest basally and a larger ventral basal bristle 0.15–0.2 from base. T 1 0.8X length of F 1, almost linear, usual series of minute adpressed denticles and outstanding hairs ventrally; at2 as long as at3. Legs otherwise with only short hairs and bristles.

Wing: with simple venation, veins brown; cell cup closed but A 1 faint, especially basally.

Abdomen: brownish to black, tergites with scattered fine black setae on disc, stronger on posterior margins, especially on tergite 5; sternites shorter haired. Terminalia ( Fig. 55 ) reflexed dorsally, hypandrium not fused with epandrium; epandrial lobes separate, slightly overlapping hypandrium below, bearing conspicuous yellowish posteroapical setae; hypandrium inverted heart-shaped; subepandrial process rather short; postgonite large, broad with flattened crescent shaped apical lobe; cerci free, at least apically, distal margin irregular bearing distinct yellowish setae and a narrow finger-like process laterally; phallus rather broad, strongly curved near base.

Description. Female: differing from male as follows. Antenna with arista longer, fully 1.5X as long as postpedicel; lpo yellowish, longer (almost as long as ocl); dc1 larger, 0.5X length of dc2. Terminal papillae slender with long fine pale hairs.

Comments. C. rangopango is a distinctive small dark species with two rows of setae beneath F 1 and all femora distinctly darkened with paler tips. It is known from just three widely distant forest and forest edge localities of the two major islands of New Zealand with capture dates from September to January. There are small differences in the male terminalia of the Urewera specimens (the apical lobe of postgonite being rather less pointed for example) but there is little doubt that they are the same species.

Figures

FIGURES 49–55. Male genitalia of Chelipoda spp. 49. C. oblata Collin, posterior view. 50. C. oblinita Collin, posterior view. 51–53. C. otiraensis (Miller). 51. posterior view. 52. anterior view. 53. apex of postgonite. 54. C. puhihiroa sp. n. apex of postgonite. 55. C. rangopango sp. n. posterior view,