Hebecnema fumosa (Meigen)

Michelsen, Verner, 2019, Macaronesian Muscidae (Diptera). I. The genus Hebecnema Schnabl with description of a new Canarian endemic species and a review of the European fauna, Zootaxa 4706 (2), pp. 332-348 : 338

Hebecnema fumosa (Meigen)

( Figs 3 A‒D)

Hydrophoria fumosa (Meigen) ; Macquart in Webb & Berthelot, 1839: 115 (Canary Is.).

Hebecnema fumosa (Meigen) ; Becker 1908 a: 95 (Gran Canaria, Tenerife, La Palma); Becker 1908b: 194 (Madeira); Frey 1935: 134 (Gran Canaria, Tenerife); Tiensuu 1938: 8 (Madeira); Frey 1944: 104 (Azores); Frey 1949: 44 (Madeira); Ringdahl 1960: 98 (Azores); Kirchberg 1967 (Hierro); Santos Abréu 1976: 73 (Canary Is.); Báez 1977: 18, figs 1, 5, 6 (Gran Canaria, Tenerife, La Gomera); Chandler 1979 (La Palma).

“ Hebecnema umbratica (Meigen) ”; Séguy 1936: 23 (Azores); Santos Abréu 1976: 77, fig. 9 (Tenerife, La Palma). Misidentifications.

Material examined. [ MZH, NHMD]: CANARY ISLANDS: GRAN CANARIA: Supra Realejo alto, 1♀ 17.v.1947 (H. Lindberg); Las Lagunetas, 1♀ 1.iv.1949 (H. Lindberg). TENERIFE: 2♂ Valle de Segovia, 500m, 13.iv.1950 (H. Lindberg); Puerto de la Cruz, 1♂ 2‒4.ii.1949 (H. Lindberg), 1♀ 16.xii. 1965 (E. Thuneberg); Santa Ursula, 1♂ 3.ii.1949 (H. Lindberg); Santa Ursula, La Quinta, 1♀ 30.iii.1993 (M. Koponen); Silos, 1♀ 12.ii.1949 (H. Lin- dberg); Los Carrizales, 28°19´N 16°51´W, c. 645m, 1♀ 10.ii.2008 (V. Michelsen). LA GOMERA: Lomito Frago- so, 28°07´N 17°10´W, 220‒250m, 2♂ 19‒24.i.2019 (V. Michelsen). LA PALMA: Las Tricias, 28°47´N 17°58´W, 660m, 1♂ 11‒16.i.2013 (V. Michelsen). MADEIRA: Quiemadas, 1♀ 14‒16.v. 1959 (S. Panelius); Serra de Agua, 1♂ 20‒21.iv.1959 (S. Panelius); Ribeira Brava, 2♂ 20.v.1959 (H. Lindberg). AZORES: Pico: Madalena, 1♀ 6‒ 9.vii.1938 (R. Storå). Further European and extralimital material in NHMD from: France ( Corsica), Greece, Italy, Portugal Spain, Sweden, Switzerland and India (Kashmir), Morocco, Tunisia and Turkey.

Diagnosis. Parafacials dull brownish grey in anterodorsal view. Thorax and abdomen with brownish grey pruinosity; female mesonotum with four distinct dark stripes in posterior view; male abdomen uniformly light brownish grey pruinose without traces of a median dark stripe; female abdomen grey pruinose with fine dark reflections and a distinct median dark stripe reaching hind margin of tergite IV. Male wing and calypteres with more or less distinct brown clouding. Halter yellow. Legs brownish black with ochre brown to pale ochre yellow tibiae. Male eye with long, moderately dense brownish pile; female eye with distinct but very short pile. Male head in profile with middle part of parafacial concealed behind the eye. Distance between rows of fine presutural acrostichal setae almost twice their distance to adjacent dorsocentral rows; setulae between acrostichal rows arranged in three or four irregular rows. Vein C bare on dorsal surface. Hind tibia with 1‒2 av setae. Male sternite V, surstylus and cerci ( Figs 3 A‒D) not clearly different from those parts in H. umbratica ( Figs 4 A‒D). Ovipositor of the Hebecnema umbratica type ( Figs 8A, B ).

Remarks. The present species belongs to the Hebecnema umbratica species group. It is evidently very closely related to H. umbratica as judged from the practically identical male and female terminalia. It is on average larger, with a wing length reaching 6.3 mm and readily recognized in both sexes by colour differences. Medium to large individuals of both sexes frequently with a few setulae on sternite I; this I have elsewhere only seen in one exceptionally large individual of H. umbratica .

Macaronesian distribution. Canary Islands: Gran Canaria, Tenerife, La Gomera, Hierro, La Palma ; Madeira; Azores.

Figures

FIGURE 3. Hebecnema fumosa (Meigen). Male terminalia. A, B. Sternite V in ventral and lateral views. C, D. Cerci, surstyli and epandrium in posterior and lateral views.

FIGURE 4. Hebecnema umbratica (Meigen). Male terminalia. A, B. Sternite V in ventral and lateral views. C, D. Cerci, surstyli and epandrium in posterior and lateral views.

FIGURE 8. Hebecnema spp. Ovipositor, left half only, except for sternites VI and VII. A, B. H. umbratica (Meigen). C. H. anthracinella sp. nov. Abbreviations: Ce = cercus; Ep = epiproct; Hy = hypoproct; St = sternite; T = tergite.